There is abundant detritus formation in this family; the outer layers of
the cortex are continually being sloughed, the dead tissues lying on the
upper surface as a dark gelatinous layer, continuous or in small patches.
On the under surface the cast-off cortex gathers into a loose confused
mass of dead tissues.
Asci 8-spored.
Spores mostly simple (disc gyrose) 1. Gyrophora Ach.
Spores 1-septate 2. *Dermatiscum Nyl.
Asci 1-2-spored.
Spores muriform 3. Umbilicaria Hoffm.
XXXV. _ACAROSPORACEAE_
Thallus foliose, squamulose or crustaceous, sometimes scarcely developed.
Algal cells Protococcaceae.
Into this family Zahlbruckner has gathered the genera in which the asci
are many-spored, as he considers that a character of great importance in
determining relationship, but he has in doing so overlooked other very
great differences. The fruit-bodies are round and completely enclosed
in a thalline wall in _Thelocarpon_, which has however no perithecial
wall. They have a proper margin only (lecideine) in _Biatorella_, and a
thalline margin (lecanorine) in the remaining genera. In _Acarospora_ the
apothecia are sunk in the thallus. Stirton’s genus _Cryptothecia_[1053]
is allied to _Thelocarpon_ in the fruit-formation, but the basal thallus
is well developed and the spores are few in number and variously divided.
Thallus none.
Apothecia (or perithecia) in thalline
warts 1. Thelocarpon Nyl.
Thallus crustaceous.
Apothecia lecideine; spores simple 2. Biatorella Th. Fr.
Apothecia lecanorine; spores septate 3. *Maronea Massal.
Thallus of small squamules 4. Acarospora Massal.
Thallus almost foliose, attached
centrally 5. *Glypholecia Nyl.
XXXVI. _EPHEBACEAE_
A family of very simple structure either filamentous, foliose or
crustaceous. The algal cells which give a dark colour to the thallus
are _Stigonema_ or _Scytonema_, members of the blue-green Myxophyceae,
and consist of minute simple or branched filaments—single cell-rows in
_Scytonema_, compound in _Stigonema_.
In some of the genera the lichen hyphae travel within the gelatinous
sheath of the filaments, both algae and hyphae increasing by apical
growth so that filaments many times the length of the alga are formed as
in _Ephebe_. In others the filaments scarcely increase beyond the normal
size of the alga as in _Thermutis (Gonionema)_; or the gelatinous algal
cells may be distributed in a stratum of hyphae.
The apothecia are minute and almost closed; they may be embedded in
swellings of the thallus, or are more or less superficial. The spores are
rather small, colourless and simple or 1-septate.
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