According to other histologists, the comparison of protoplasm to a
spongy mass does not give the most exact idea, and, in particular, it
does not furnish the most general idea. It would be far better to say
that the protoplasm possesses the structure of foam or lather. As was
seen by Kunstler in 1880, a comparison with some familiar objects gives
the best idea. Nothing could be more like protoplasm physically than
the culinary preparation known as _sauce mayonnaise_, made with the aid
of oil and a liquid with which oil does not mix. Emulsions of this kind
were made artificially by Bütschli. He noted that these preparations
mimicked all the aspects of cellular protoplasm. Thus, in the living
cell there is a mixture of two liquids, non-miscible and of unequal
fluidity. This mixture gives rise to the formation of little cells. The
more consistent substance forms their supporting framework (Leydig’s
spongioplasm), while the other, which is more fluid, fills its interior
(hyaloplasm).
However that may be, whether the primitive organization of the cellular
protoplasm be that of a sponge, as is asserted by Leydig, or that of
a _sauce mayonnaise_, as is claimed by Bütschli and Kunstler, the
complexity does not rest there. Further recourse must be made to
analysis. Just as the tissue of a sponge, when torn, shows the fibres
which constitute it, so the spongioplasm, the parietal substance,
is exhibited as formed of a tangle of fibrils, or better still, of
filaments or ribbons (in Greek, _mitome_), which are called _chromatic
filaments_, because they are deeply stained when the cell is plunged
into aniline dye. In each of these filaments, the substance of which
is called chromatin, the devices of microscopic examination enable
us to discover a series of granulations like beads on a string, the
_microsomes_ or bioblasts, connected one with the other by a sort of
cement, Schwartz’s _linin_, which is a kind of nuclein.
And let us add, to complete this summary of the constitution of
cellular protoplasm, that it presents, at any rate at a certain moment,
a remarkable organ, the _centrosome_, which plays an important part
in cellular division. Its pre-existence is not certain. Some writers
make it issue from the nucleus. At the moment of cellular division it
appears like a compressed mass of granulations, which may be deeply
stained. Around it is seen a clear unstainable zone, called the
attraction-sphere; and finally, beyond this is a crown of striæ, which
diverge like the rays of a halo—_i.e._, the _aster_. In conclusion,
there are yet in the cellular body three kinds of non-essential bodies:
the vacuoles, the leucites, and various inclusions. The _vacuoles_ are
cavities, some inert, some contractile; the _leucites_ are organs for
the manufacture of particular substances; the _inclusions_ are the
manufactured products, or wastes.
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