either in sterility or variation—how much of the work of originating
species is left for natural selection?—which, as Mr. Darwin admits
(“Natural Selection,” p. 63, ed. 1876), does not _induce variability_,
but “implies only the preservation of _such variations as arise_, and are
beneficial to the being under its conditions of life?” An important part
assuredly, and one which we can never sufficiently thank Mr. Darwin for
having put so forcibly before us, but an indirect part only, like the
part played by time and space, and not, I think, the one which Mr. Darwin
would assign to it.
Mr. Darwin himself has admitted that in the earlier editions of his
“Origin of Species” he “underrated, as it now seems probable, the
frequency and importance of modifications due to spontaneous
variability.” And this involves the having over-rated the action of
“natural selection” as an agent in the evolution of species. But one
gathers that he still believes the accumulation of small and fortuitous
variations through the agency of “natural selection” to be the main cause
of the present divergencies of structure and instinct. I do not,
however, think that Mr. Darwin is clear about his own meaning. I think
the prominence given to “natural selection” in connection with the
“origin of species” has led him, in spite of himself, and in spite of his
being on his guard (as is clearly shown by the paragraph on page 63
“Natural Selection,” above referred to), to regard “natural selection” as
in some way accounting for variation, just as the use of the dangerous
word “spontaneous,”—though he is so often on his guard against it, and so
frequently prefaces it with the words “so-called,”—would seem to have led
him into very serious confusion of thought in the passage quoted at the
beginning of this paragraph.
For after saying that he had underrated “the frequency and importance of
modifications due to spontaneous variability,” he continues, “but it is
impossible to attribute to this cause the innumerable structures which
are so well adapted to the habits of life of each species.” That is to
say, it is impossible to attribute these innumerable structures to
spontaneous variability.
What _is_ spontaneous variability?
Clearly, from his preceding paragraph, Mr. Darwin means only “so-called
spontaneous variations,” such as “the appearance of a moss-rose on a
common rose, or of a nectarine on a peach-tree,” which he gives as good
examples of so-called spontaneous variation.
Public-domain text, read in full here on John Shaqi.
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