The oak family can be traced back to the Miocene period, and consequently
enjoys quite as high an antiquity as the sequoias. Professor Gray, in
speaking of the _Quercus robur_ and its probable origin, says that it is
"traceable in Europe up to the commencement of the present epoch, looks
eastward, and far into the past on far-distant shores." By "far-distant
shores," he undoubtedly means Northwest America, where its remotest
descendants still flourish. But that these trees should have waded the
Pacific, or sent their acorns on a voyage of discovery after new habitats
on the Asiatic coast, is hardly more probable than Jason's voyage after
the golden fleece, in any other than a highly figurative sense. The
spontaneous appearance of a forest of oaks on the eastern shores of Asia
was just as probable, under favoring conditions--though occurring
subsequently to the time of their appearance on this continent--as that of
the miniature forests of "samphire," or small saline plants, which
spontaneously made their appearance about the salt-works of Syracuse, when
conditions actually favored. The high antiquity of the oak makes no
difference in respect to the principle of dispersion, since geographical
conditions are what govern, and not the theoretical considerations of the
speculative botanist.
Mr. A. R. Wallace's formula concerning the origin of species, that they
"have come into existence coincident both in time and place with
preëxisting closely-allied species," may or may not be true so far as
individual localization is concerned. But it proves nothing in the way of
original progeny, nor can we, by any actual data before us, satisfactorily
determine, under this formula, which of the two closely-allied species
preceded the other. If they came coincidently, both in time and place,
their existence must have been concurrent, not separated by preëxistence.
The formula may be true to this extent, that the conditions favoring the
appearance of one species may have equally favored what we call a
closely-allied species. But even in this case, the material sequence is
lost, and we have nothing to express a relationship as from parent to
progeny. For, however restricted as to localization, each species
preserves its own characteristics, the similarities always being less than
the dissimilarities. These, and other equally conclusive facts of
observation, led Professor Agassiz to question any necessary genetic
connection between the different species, or between even the same
species, in widely-separated localities; his idea being precisely that
advanced by us in connection with the Bible genesis, that localization
depended on geographical conditions, not on the migration of plants or the
dispersion of seeds.
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