Life Movements in Plants, Volume IBose, Jagadis Chandra
Science
Life Movements in Plants, Volume I
Bose, Jagadis Chandra
Growth (Plants); Plants -- Irritability and movements
The practical difficulties met with in carrying out this idea are
very numerous. It will be understood that just as the imperceptible
movement is highly magnified by the compound system of levers, the
various errors and difficulties are likely to be magnified in the
same proportion. The principal difficulties met with were due: (1) to
the weight of the compound lever which exerted a great tension on the
growing plant, (2) to the yielding of flexible connections by which
the plant was attached to the first lever, and the first lever to the
second, and (3) to the friction at the fulcrums.
_Weight of the Lever._--As the first lever is to exert a pull on
the second, it has to be made rigid. The second lever serves as an
index, and can therefore be made of fine glass fibre. The securing of
rigidity of the first lever entails large cross section and consequent
weight, which exerts considerable tension on the plant. Excessive
tension greatly modifies growth; even the weight of the index used
in self-recording auxanometers is found to modify the normal rate of
growth. The weight of the levers introduces an additional difficulty in
the increased friction at the fulcrums, on account of which there is an
obstruction of the free movement of the recording arm of the lever. The
conditions essential for overcoming these difficulties therefore are:
(1) construction of a very light lever possessing sufficient rigidity,
and (2) arranging the levers in such a way that the tension on the
plant may be reduced to any extent, or even eliminated.
I found in _navaldum_, an alloy of aluminium, a light material
possessing sufficient rigidity. The first lever is constructed out of a
thin narrow sheet 25 cm. in length; it has, as explained before, to be
fairly rigid in order to exert a pull on the second without undergoing
any bending; this rigidity is secured by giving the thin narrow plate
of the lever a T-shape. The first lever balances, to a certain extent,
the second. Finer adjustments are made by means of an adjustable
counterpoise B, at the end of the levers. By this means the tension on
the plant can be greatly reduced; or a constant tension may be exerted
by means of a weight T (Fig. 56). In my later type of the apparatus
the plant connection is made to the right, instead of the left side of
the first fulcrum. This gives certain practical advantages. The second
lever is then made practically to balance the first, only a very slight
weight being necessary for exact counterpoise. The reduction of total
weight thus secured reduces materially the friction at the fulcrum with
great enhancement of efficiency of the apparatus.
[Illustration: FIG. 56.--Compound lever. P, plant attached to short arm
of lever L; T, weight exerting tension; C, connecting link; L′, second
lever with bent tip for record; B, B, balancing counterpoise. Fork F,
carries at its side two conical agate cups, on which lever rests by two
pin-points. (From a photograph.)]
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