Life Movements in Plants, Volume IIBose, Jagadis Chandra
Science
Life Movements in Plants, Volume II
Bose, Jagadis Chandra
Growth (Plants); Plants -- Irritability and movements
_Photo-geotropic balance: Experiment 195._--I shall here describe in
detail the procedure for obtaining an exact balance. A parallel beam of
light from a small arc lamp is reflected by means of an inclined mirror,
so as to act on the pulvinus below. An iris diaphragm regulates the
intensity of incident light. The first part of the curve is the record
of geotropic torsional movement. Light of a given intensity was applied
below at a point marked -L (Fig. 181); this is seen to produce an
over-balance, the phototropic effect being slightly in excess. The
intensity of incident light was continuously diminished by regulation of
the diaphragm till an exact balance was obtained as seen in the
horizontal part of the record. It is with great surprise that one comes
to realise the fact that the effect of one form of stimulus can be so
exactly balanced by that of another, so entirely different, and that the
stimulus of gravity could be measured, as it were, in candle powers of
light! After securing the balance, light was cut off, and the geotropic
torsion became renewed on the cessation of the counteracting phototropic
action.
[Illustration: FIG. 182.--Application of white light at--L in opposition
causes reversal of torsion. Red light R, is ineffective, and geotropic
torsion is restored. Reapplication of white light causes once more the
reversal of torsion.]
_Comparative balancing effects of white and red lights: Experiment
196._--White light was at first applied at -L in opposition to geotropic
movement. The intensity of light was stronger than what was necessary
for exact balance, and its effect was at first to retard and then
reverse the torsional response due to geotropism. When thus
overbalanced, red glass was interposed on the path of light at R. As the
phototropic effect of this light is feeble or absent, the geotropic
torsion became predominant as seen in the subsequent up-curve. The red
glass was next removed substituting white light at -L to act once more
in opposition; the result is seen in the final over-balance, and
reversal of torsion (Fig. 182).
[Illustration: FIG. 183.--Effect of coal gas on photo geotropic balance.
Geotropic torsion, G, is exactly balanced by opposing action of light
-L. Application of coal gas at C, at first caused enhancement of
phototropic action with resulting reversal. Prolonged application
induced depression of phototropic reaction, geotropic action thus
becoming predominant.]
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