Life Movements in Plants, Volume IIBose, Jagadis Chandra
Science
Life Movements in Plants, Volume II
Bose, Jagadis Chandra
Growth (Plants); Plants -- Irritability and movements
The question now arises whether this position of geotropic equilibrium
is permanent, or whether it undergoes modification in a definite way by
variation of temperature. I shall proceed to show that the position of
equilibrium undergoes a change in one direction by a rise, and in the
opposite direction by a fall of temperature. I shall use the term
_thermo-geotropism_ as a convenient phrase to indicate the effect of
temperature in modification of geotropic curvature and torsion.
I shall first deal with the effect of variation of temperature on
geotropic torsion. Under the continued action of stimulus of gravity the
torsion increases till it reaches a limit; for the twisted organ
resists further distortion and a balance is struck when the twisting and
untwisting forces are equal and opposite. In this state of equilibrium
the effect of an external agent, say of variation of temperature, will
bring about an upset of the balance. The torsion will be increased if
the external agent induces an enhancement of geotropic action; it will,
on the other hand, be decreased when it induces a diminished reaction.
[Illustration: FIG. 184.--Magnet M causes deflection of the needle _n
s_, suspended by a thin wire. Increase of magnetisation of M increases
deflection, while decrease of magnetisation diminishes the deflection.]
A physical analogy will make this point clear; imagine a small magnetic
needle suspended by a thin wire; the earth's directive force is supposed
to be annulled by the well known device of a compensating magnet. A
second and larger magnet M is now placed at right angles to the
suspended needle; N will repel _n_ and attract _s_, and a deflection
will be produced, the deflecting force of the magnet M being balanced by
the force of torsion of suspending wire (Fig. 184).
The state of equilibrium will however be disturbed by variation of the
magnetic force of M. It is known that a rise of temperature diminishes
magnetisation while lowering of temperature increases it. Hence the
deflecting force of the magnet will be diminished under rise of
temperature with concomitant diminution of deflection of the needle and
the torsion of the wire. Fall of temperature, on the other hand, will
cause an increase of deflection and of torsion. The physical
illustration given above will help us to understand how the
physiological effect of variation of temperature may bring about changes
in geotropic curvature and torsion.
TROPIC EQUILIBRIUM UNDER VARYING INTENSITIES OF STIMULUS.
The following experiment will show that the position of tropic
equilibrium is not fixed but subject to variation under changes of
effective stimulation.
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