“I should point out that, according to the evolutionists of the last century, improvement in the eye, as in any other organ, is mainly due to persistent, rational, employment of the organ in question, in such slightly modified manner as experience and changed surroundings may suggest. You will have observed that, according to my system, this goes for very little, and that the accumulation of fortunate accidents, irrespectively of the use that may be made of them, is by far the most important means of modification. Put more briefly still, the distinction between me and my predecessors lies in this;—my predecessors thought they knew the main normal cause or principle that underlies variation, whereas I think that there is no general principle underlying it at all, or that even if there is, we know hardly anything about it. This is my distinctive feature; there is no deception; I shall not consider the arguments of my predecessors, nor show in what respect they are insufficient; in fact, I shall say nothing whatever about them. Please to understand that I alone am in possession of the master key that can unlock the bars of the future progress of evolutionary science; so great an improvement, in fact, is my discovery that it justifies me in claiming the theory of descent generally, and I accordingly claim it. If you ask me in what my discovery consists, I reply in this;—that the variations which we are all agreed accumulate are caused—by variation. {209a} I admit that this is not telling you much about them, but it is as much as I think proper to say at present; above all things, let me caution you against thinking that there is any principle of general application underlying variation.” This would have been right. This is what Mr. Darwin would have had to have said if he had been frank with us; it is not surprising, therefore, that he should have been less frank than might have been wished. I have no doubt that many a time between 1859 and 1882, the year of his death, Mr. Darwin bitterly regretted his initial error, and would have been only too thankful to repair it, but he could only put the difference between himself and the early evolutionists clearly before his readers at the cost of seeing his own system come tumbling down like a pack of cards; this was more than he could stand, so he buried his face, ostrich-like, in the sand. I know no more pitiable figure in either literature or science.