Makers of British Botany; a collection of biographies by living botanists
History
Makers of British Botany; a collection of biographies by living botanists
Botanists -- Great Britain -- Biography; Botany -- Great Britain -- History
Comparatively little was known of the life history of any Schizomycete.
Ward therefore made a detailed and exhaustive study of that of
_Bacillus ramosus_, the _Wurzel bacillus_ of German authors, which is
common in Thames water, and bears a superficial resemblance to the
anthrax bacillus, but is innocuous. It proved convenient for study, as
it ran through its entire life history in from thirty to sixty hours
at ordinary temperature. It forms long filaments, the growth of which
Ward was able to measure under the microscope with great precision. On
plotting out his measurements he obtained a regular curve, from which
he found that, under constant conditions, the filament doubled itself
in equal times. This he called "the law of doubling." It is the same as
the so-called "law of compound interest," and leads to the expression
of the growing quantity as an exponential function of the time, so
that the time is proportional to the logarithm of that quantity. This
relation has, of course, long been familiar in chemical reactions, but,
as far as I know, Ward was the first to detect it in any vital process
in a plant. This, which was in 1895, has, I think, been overlooked.
Stefanowska has since, in 1904, obtained a logarithmic curve for the
early period of the growth of maize, which doubles its weight every ten
days, and the subject has since been pursued by Chodat and others.
In speculating on the cause of the destructive action of light on
bacteria, Ward adopted the view of his friend Elfving, that it
inhibited metabolic processes necessary to nutrition. He suggests that
the "constructed metabolites" at the moment of assimilation are in
a highly unstable condition, and liable to destruction by oxidation
promoted by light. He points to the fact that plant structures are
frequently provided with colour screens, which would cut off the
blue-violet rays and check their action in promoting the rapid
oxidation of reserve materials, and he quotes the suggestion of Elfving
that chlorophyll itself may serve as such a screen against "destructive
metabolic action in synthesis." Ward seems to have attributed little
importance to the fact that substantially the same view had long before
been put forward by Pringsheim, though received with little favour. His
own view that when red and orange predominate in the screens their
effect is protective, has since afforded a probable explanation of the
colouration of young foliage, especially in the tropics.
Public-domain text, read in full here on John Shaqi.
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