Makers of British Botany; a collection of biographies by living botanists
History
Makers of British Botany; a collection of biographies by living botanists
Botanists -- Great Britain -- Biography; Botany -- Great Britain -- History
The dedication is to George Prince of Wales, afterwards George III.
The author cannot quite avoid the style of his day, for instance: "And
as _Solomon_ the greatest and wisest of men, deigned[28] to inquire
into the nature of Plants, _from the Cedar of Lebanon, to the Hyssop
that springeth out of the wall_. So it will not, I presume, be an
unacceptable entertainment to your Royal Highness," etc.
But the real interest of the dedication is its clear statement of
his views on the nutrition of plants. He asserts that plants obtain
nourishment, not only from the earth, "but also more sublimed and
exalted food from the air, that wonderful fluid, which is of such
importance to the life of Vegetables and Animals," etc. We shall see
that his later statement is not so definite, and it is well to rescue
this downright assertion from oblivion.
His book begins with the research for which he is best known, namely
that on transpiration. He took a sunflower growing in a flower-pot,
covering the surface of the earth with a plate of thin milled lead,
and cemented it so that no vapour could pass, leaving a corked hole to
allow of the plant being watered. He did not take steps to prevent loss
through the pot, but at the end of the experiment cut off the plant,
cemented the stump and found that the "unglazed porous pot" perspired 2
ozs. in 12 hours, and for this he made due allowance.
The plant so prepared he proceeded to weigh at stated intervals. He
obtained the area of the leaves by dividing them into parcels according
to their several sizes and measuring one leaf[29] of each parcel. The
loss of water in 12 hours converted to the metric system is 1·3 c.c.
per 100 sq. cm. of leaf-surface; and this is of the same order of
magnitude as Sachs' result[30], namely 2·2 c.c. per 100 sq. cm. He
goes on to measure the surface of the roots[31] and to estimate the
rate of absorption per area. The calculation is of no value, since
he did not know how small a part of the roots is absorbent, nor how
enormously the surface of that part is increased by the presence of
root-hairs. He goes on to estimate the rate of the flow of water up
the stem; this would be 34 cubic inches in 12 hours if the stem (which
was one square inch in section) were a hollow tube. He then allowed a
sunflower stem to wither and to become completely dry, and found that
it had lost ¾ of its weight, and assuming that the ¼ of the "solid
parts" left was useless for the transmission of water he increases his
34 by ⅓ and gives 45⅓ cubic inches in 12 hours as the rate. But
the solid matter which he neglected contained the vessels and he would
have been nearer to the truth had he corrected his figures on this
basis. The simplest plan is to compare his results with those obtained
by Sachs[32] in allowing plants to absorb solutions of lithium-salts.
If the flow takes place through conduits equivalent to a quarter of a
square inch in area, the fluid will rise in 12 hours to a height of
Public-domain text, read in full here on John Shaqi.
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