Professor Boyd Dawkins attempts to reduce the difficulty by supposing that the Glacial epoch was marked by the occurrence of minor periods of climatic variation, during which, in comparatively short periods, the isothermal lines vibrated from north to south, and _vice versa_. In this view the southern species gradually crowded upon the northern during the periods of climatic amelioration, until they reached their limit in central England, and then in turn, as the climate became more rigorous, slowly retreated before the pressure of their northern competitors. Meanwhile the hyena sallied forth from his various caves, over this region, at one time of the year to feed upon the reindeer, and at another time of the year upon the flesh of the hippopotamus, in both cases dragging their bones with him to his sheltered retreat in the limestone caverns[DN] which he shared at intervals with palæolithic man. [Footnote DN: Early Man in Britain, p. 114.] The theory of Mr. James Geikie is that the period, while one of great precipitation, was characterised by a climate of comparatively even temperature, in which there was not so great a difference as now between the winters and the summers, the winters not being so cold and the summers not so hot as at present. This is substantially the condition of things in southern Alaska at the present time, where extensive glaciers come down to the sea-level, even though the thermometer at Sitka rarely goes below zero (Fahrenheit). It is, therefore, easy to conceive that if there were extensive plains bordering the Alaskan archipelago, so as to furnish ranging grounds for more southern species, the animals of the north and the animals of the south might partially occupy the same belt of territory, and their bones become mingled in the same river deposits. In order to clear the way for either of these hypotheses to account for the mingling of arctic and torrid species characteristic of the period under consideration in Europe, we must probably suppose such an elevation of the region to the south as to afford land connection between Europe and Africa. This would be furnished by only a moderate amount of elevation across the Strait of Gibraltar and from the south of Italy to the opposite shore in Africa; and there are many indications, in the distribution of species, of the existence in late geological times of such connection.