Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
The _Telephone_ case may have been of the same nature; though, as we
have seen above, this pea is peculiar in its colour-heredity and may
quite well have followed a different rule in shape also. As stated
before, the wrinkled offspring were not cultivated after the third
year, but the _round_ seeds are said to have still given some wrinkleds
in the eighth year after the cross, as would be expected in a simple
Mendelian case.
(_b_) _Tschermak’s cases._ The cases Professor Weldon quotes from
Tschermak all relate to crosses with _Telephone_ again, and this fact
taken with the certainty that the colour-heredity of _Telephone_ is
abnormal makes it fairly clear that there is here something of a really
exceptional character. What the real nature of the exception is, and
how far it is to be taken as contradicting the “law of dominance,” is
quite another matter.
3. _Other phenomena, especially regarding seed-shapes, in the case of
“grey” peas. Modern evidence._ Professor Weldon quotes from Tschermak
the interesting facts about the “grey” pea, _Graue Riesen_, but does
not attempt to elucidate them. He is not on very safe ground in
adducing these phenomena as conflicting with the “law of dominance.”
Let us see whither we are led if we consider these cases. On p. 124
I mentioned that the classes round and wrinkled do not properly hold
if we try to extend them to large-seeded sorts, and that these cases
require separate consideration. In many of such peas, which usually
belong either to the classes of sugar-peas (_mange-touts_) or “grey”
peas (with coloured flowers), the seeds would be rather described as
irregularly indented, lumpy or stony[94], than by any use of the terms
round or wrinkled. One sugar-pea (_Debarbieux_) which I have used
has large flattish, smooth, yellow seeds with white skins, and this
also in its crossings follows the rules about to be described for the
large-seeded “grey” peas.
[94] Gärtner’s _macrospermum_ was evidently one of these, though
from the further account (p. 498) it was probably more wrinkled.
There are of course _mange-touts_ which have perfectly round seeds.
Mendel himself showed that the _mange-tout_ character, the soft
constricted pod, was transferable. There are also _mange-touts_
with fully wrinkled seeds and “grey” peas with small seeds (see
Vilmorin-Andrieux, _Plantes Potagères_, 1883).
In the large “grey” peas the most conspicuous feature is the seed-coat,
which is grey, brownish, or of a bright reddish colour. Such seed-coats
are often speckled with purple, and on boiling these seed-coats turn
dark brown. They are in fact the very peas used by Mendel in making up
his third pair of characters. Regarding them Professor Weldon, stating
they may be considered separately, writes as follows:--
Public-domain text, read in full here on John Shaqi.
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