Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
In 1867 I sowed these seeds, and all five produced tall purple-flowered
purplish-stemmed plants[106], and the seeds, with few exceptions,
had all maple or brownish-streaked envelopes of various shades; the
remainder had entirely violet or deep purple-coloured envelopes[107]:
in shape the peas were partly indented; but a few were round[108].
Some of the plants ripened off earlier than the “maple,” which, in
comparison with “Ringleader,” is a late variety; and although the
pods were in many instances partially abortive, the produce was very
large[109].
[106] Tall heterozygotes, with normal dominance of purple flowers.
[107] Here we see dominance of the _pigmented_ seed-coat as a
maternal character over _white_ seed-coat. The colours of the
seed-coats are described as essentially two: maple or brown-streaked,
and violet, the latter being a small minority. As the sequel shows,
the latter are heterozygotes, not breeding true. Now Mendel found,
and the fact has been confirmed both by Correns and myself, that
crossing a grey pea which is capable of producing purple leads to
such production as a form of xenia.
We have here therefore in the purple seeds the union of dissimilar
gametes, with production of xenia. But as the brown-streaked
seeds are also in part heterozygous, the splitting of a compound
allelomorph has probably taken place, though without precise
statistics and allotment of offspring among the several seeds the
point is uncertain. The colour of seed-coats in “grey” peas and
probably “maples” also is, as was stated on p. 150, sensitive to
conditions, but the whole difference between “maples” and purple is
too much to attribute safely to such irregularity. “Maple” is the
word used to describe certain seed-coats which are pigmented with
intricate brown mottlings on a paler buff ground. In French they are
_perdrix_.
[108] This is not, as it stands, explicable. It seems from this point
and also from what follows that if the account is truly given, some
of the plants may have been mosaic with segregation of characters in
particular flowers; but see subsequent note.
[109] As, commonly, in heterozygotes when fertile.
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