Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
From the above I gather that the white-flowered white-seeded pea is
(if I may use the term) an original variety well fixed and distinct
entirely from the maple, that the two do not thoroughly intermingle
(for whenever the white flower crops out, the plant and its parts all
appear to follow exactly the characters of the white pea), and that the
maple is a cross-bred variety which has become somewhat permanent and
would seem to include amongst its ancestors one or more bearing seeds
either altogether or partly violet- or purple-coloured; for although
this colour does not appear on the seed of the “maple,” it is very
potent in the variety, and appears in many parts of the plant and its
offspring from cross-fertilised flowers, sometimes on the external
surface or at the sutures of the pods of the latter, at others on
the seeds and stems, and very frequently on the seeds; and whenever
it shows itself on any part of the plant, the flowers are invariably
purple. My deductions have been confirmed by intercrosses effected
between the various white-, blue-, some singularly bright green-seeded
peas which I have selected, and the maple- and purple-podded and the
purple-flowered sugar peas, and by reversing those crosses.
I have also deduced from my experiments, in accordance with the
conclusions of the late Mr Knight and others, that the colours of the
envelopes of the seeds of peas immediately resulting from a cross are
never changed[121]. I find, however, that the colour and probably the
substance of the cotyledons are sometimes, but not always, changed
by the cross fertilisation of two different varieties; and I do not
agree with Mr Knight that the form and size of the seeds produced are
unaltered[122]; for I have on more than one occasion observed that
the cotyledons in the seeds directly resulting from a cross of a blue
wrinkled pea fertilised by the pollen of a white round variety have
been of a greenish-white colour[123], and the seeds nearly round[124]
and larger or smaller according as there may have been a difference in
the size of the seeds of the two varieties[125].
[121] The nature of this mistake is now clear; for as stated above
xenia is only likely to occur when the maternal seed-coat is
pigmented. The violet coats in this experiment are themselves cases
of xenia.
[122] Knight, it was seen, crossed round ♀ × indented ♂ and
consequently got no change of form.
[123] Cotyledons seen through coat.
[124] Ordinary dominance of round.
[125] This is an extraordinary statement to be given as a general
truth. There are sometimes indications of this kind, but certainly
the facts are not usually as here stated.
Public-domain text, read in full here on John Shaqi.
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