Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
Professor Weldon declares that Laxton’s “experience was altogether
different from that of Mendel.” The reader will bear in mind that when
Laxton speaks of fixing a variety he is not thinking particularly of
seed-characters, but of all the complex characters, fertility, size,
flavour, season of maturity, hardiness, etc., which go to make a
serviceable pea. Considered carefully, Laxton’s testimony is so closely
in accord with Mendelian expectation that I can imagine no chance
description in non-Mendelian language more accurately stating the
phenomena.
Here we are told in unmistakable terms the breaking up of the original
combination of characters on crossing, their re-arrangement, that
at the fourth or fifth generation the possibilities of sporting
[sub-division of compound allelomorphs and re-combinations of them?]
are exhausted, that there are then definite forms which if selected
are thenceforth fixed [produced by union of similar gametes?]
that it takes longer to select some forms [dominants?] than others
[recessives?], that there may be “mule” forms[144] or forms which
cannot be fixed at all[145] [produced by union of dissimilar gametes?].
[144] For instance the _talls_ produced by crossing _dwarfs_ are such
“mules.” Tschermak found in certain cases distinct increase in height
in such a case, though not always (p. 531).
[145] “The remarkably fine but unfixable pea _Evolution_.” Laxton, p.
37.
But Laxton tells us more than this. He shows us that numbers of
varieties may be obtained--hundreds--“incalculable numbers.” Here
too if Professor Weldon had followed Mendel with even moderate care
he would have found the secret. For in dealing with the crosses of
_Phaseolus_ Mendel clearly forecasts the conception of _compound
characters themselves again consisting of definite units_, all of which
may be separated and re-combined in the possible combinations, laying
for us the foundation of the new science of Analytical Biology.
How did Professor Weldon, after reading Mendel, fail to perceive these
principles permeating Laxton’s facts? Laxton must have seen the very
things that Mendel saw, and had he with his other gifts combined that
penetration which detects a great principle hidden in the thin mist of
“exceptions,” we should have been able to claim for him that honour
which must ever be Mendel’s in the history of discovery.
Public-domain text, read in full here on John Shaqi.
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