Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
If there were no ancestry, there would be no posterity. But to answer
the question _why_ certain of the posterity depart from the rule which
others follow, we must know, not the ancestry, but how it came about
_either_ that at a certain moment a certain gamete divided from its
fellows in a special and unwonted fashion; _or_, though the words
are in part tautological, the reason why the union of two particular
gametes in fertilisation took place in such a way that gametes having
new specific properties resulted[161]. No one yet knows how to use the
facts of ancestry for the elucidation of these questions, or how to get
from them a truth more precise than that contained in the statement
that a diversity of specific consequences (in heredity) may follow an
apparently single specific disturbance. Rarely even can we see so much.
The appeal to ancestry, as introduced by Professor Weldon, masks the
difficulty he dare not face.
[161] May there be a connection between the extraordinary fertility
and success of the _Telephone_ group of peas, and the peculiar
frequency of a blended or mosaic condition of their allelomorphs?
The conjecture may be wild, but it is not impossible that the two
phenomena may be interdependent.
In other words, it is the _cause of variation_ we are here seeking.
To attack that problem no one has yet shown the way. Knowledge of a
different order is wanted for that task; and a compilation of ancestry,
valuable as the exercise may be, does not provide that particular kind
of knowledge.
Of course when once we have discovered by experiment that--say,
_Telephone_--manifests a peculiar behaviour in heredity, we can perhaps
make certain forecasts regarding it with fair correctness; but that
any given race or individual will behave in such a way, is a fact not
deducible from its ancestry, for the simple reason that organisms
of identical ancestry may behave in wholly distinct, though often
definite, ways.
It is from this hitherto hopeless paradox that Mendel has begun at last
to deliver us. The appeal to ancestry is a substitution of darkness for
light.
VII. THE QUESTION OF ABSOLUTE PURITY OF GERM-CELLS.
But let us go back to the cases of defective “purity” and consider how
the laws of ancestry stand in regard to them. It appears from the facts
almost certain that purity may sometimes be wanting in a character
which elsewhere usually manifests it.
Here we approach a question of greater theoretical consequence to the
right apprehension of the part borne by Mendelian principles in the
physiology of heredity. We have to consider the question whether the
purity of the gametes in respect of one or other antagonistic character
is or is likely to be in case of _any_ given character a _universal_
truth? The answer is unquestionably--No--but for reasons in which
“ancestry” plays no part[162].
[162] This discussion leaves “false hybridism” for separate
consideration.
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