Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
I am quite willing to suppose, with Professor Weldon (p. 248), that the
pea _Stratagem_ may, as he suggests, be such a case. I am even willing
to accept provisionally as probable that when two gametes, themselves
of mosaic or blended character, meet together in fertilisation, they
are more likely to produce gametes of mosaic or blended character than
of simply discontinuous character. Among Messrs Sutton’s Primulas
there are at least two striking cases of “flaked” or “bizarre” unions
of bright colours and white which reproduce themselves by seed with
fair constancy, though Mendelian purity in respect of these colours
is elsewhere common in the varieties (I suspect mosaics of “false
hybridism” among allelomorphs in some of these cases). Similarly Galton
has shown that though children having one light-eyed and one dark-eyed
parent generally have eyes either light or dark, the comparatively rare
medium eye-coloured persons when they mate together frequently produce
children with medium eye-colour.
In this connection it may be worth while to allude to a point of some
practical consequence. We know that when pure dominant--say yellow--is
crossed with pure recessive--say green--the dominance of yellow is
seen; and we have every reason to believe this rule generally (_not_
universally) true for pure varieties of peas. But we notice that in
the case of a form like the pea, depending on human selection for its
existence, it might be possible in a few years for the races with pure
seed characters to be practically supplanted by the “mosaicized” races
like the _Telephone_ group, if the market found in these latter some
specially serviceable quality. In the maincrop peas I suspect this very
process is taking place[163]. After such a revolution it might be
possible for a future experimenter to conclude that _Pisum sativum_ was
by nature a “mosaicized” species in these respects, though the mosaic
character may have arisen once in a seed or two as an exceptional
phenomenon. When the same reasoning is extended to wild forms depending
on other agencies for selection, some interesting conclusions may be
reached.
[163] Another practical point of the same nature arises from the
great variability which these peas manifest in plant- as well as
seed-characters. Mr Hurst of Burbage tells me that in _e.g._ _William
the First_, a pea very variable in seed-characters also, tall plants
may be so common that they have to be rogued out even when the
variety is grown for the vegetable market, and that the same is true
of several such varieties. It seems by no means improbable that it is
by such roguing that the unstable mosaic or blend-form is preserved.
In a thoroughly stable variety such as _Ne Plus Ultra_ roguing is
hardly necessary even for the seed-market.
Public-domain text, read in full here on John Shaqi.
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