Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
I have now shown how the mosaic and blend-forms are to be regarded in
the light of the Mendelian principle. What has Professor Weldon to say
in reference to them? His suggestion is definite enough--that a study
of ancestry will explain the facts: _how_, we are not told.
In speaking of the need of study of the characters of the _race_ he
is much nearer the mark, but when he adds “that is their ancestry,”
he goes wide again. When _Telephone_ does not truly divide the
antagonistic characters among its germ-cells this fact is in nowise
simply traceable to its having originated in a cross--a history it
shares with almost all the peas in the market--but to its own peculiar
nature. In such a case imperfect dominance need not surprise us.
What we need in all these phenomena is a knowledge of the properties
of each race, or variety, as we call it in peas. We must, as I have
often pleaded, study the properties of each form no otherwise than the
chemist does the properties of his substances, and thus only can we
hope to work our way through these phenomena. _Ancestry_ holds no key
to these facts; for the same ancestry is common to own brothers and
sisters endowed with dissimilar properties and producing dissimilar
posterity. To the knowledge of the properties of each form and the laws
which it obeys there are no short cuts. We have no periodic law to
guide us. Each case must as yet be separately worked out.
We can scarcely avoid mention of a further category of phenomena
that are certain to be adduced in opposition to the general truth
of the purity of the extracted forms. It is a fact well known to
breeders that a highly-bred stock may, unless selections be continued,
“degenerate.” This has often been insisted on in regard to peas. I
have been told of specific cases by Messrs Sutton and Sons, instances
which could be multiplied. Surely, will reply the supporters of the
theory of Ancestry, this is simply impurity in the extracted stocks
manifesting itself at last. Such a conclusion by no means follows, and
the proof that it is inapplicable is obtained from the fact that the
“degeneration,” or variation as we should rather call it, need not
lead to the production of any proximate ancestor of the selected stock
at all, but immediately to a new form, or to one much more remote--in
the case of some high class peas, _e.g._, to the form which Mr Sutton
describes as “vetch-like,” with short pods, and a very few small round
seeds, two or three in a pod. Such plants are recognized by their
appearance and are rigorously hoed out every year before seeding.
Public-domain text, read in full here on John Shaqi.
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