Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
In the process of fertilisation each pollen form unites on an average
equally often with each egg cell form, so that each of the four pollen
cells _AB_ unites once with one of the forms of egg cell _AB_, _Ab_,
_aB_, _ab_. In precisely the same way the rest of the pollen cells
of the forms _Ab_, _aB_, _ab_ unite with all the other egg cells. We
obtain therefore--
_AB_/_AB_ + _AB_/_Ab_ + _AB_/_aB_ + _AB_/_ab_ + _Ab_/_AB_ + _Ab_/_Ab_ +
_Ab_/_aB_ + _Ab_/_ab_ + _aB_/_AB_ + _aB_/_Ab_ + _aB_/_aB_ + _aB_/_ab_ +
_ab_/_AB_ + _ab_/_Ab_ + _ab_/_aB_ + _ab_/_ab_,
or
_AB_ + _ABb_ + _AaB_ + _AaBb_ + _ABb_ + _Ab_ + _AaBb_ + _Aab_ + _AaB_ +
_AaBb_ + _aB_ + _aBb_ + _AaBb_ + _Aab_ + _aBb_ + _ab_ = _AB_ + _Ab_ +
_aB_ + _ab_ + 2_ABb_ + 2_aBb_ + 2_AaB_ + 2_Aab_ + 4_AaBb_[42].
[42] [In the original the sign of equality (=) is here represented by
+, evidently a misprint.]
In precisely similar fashion is the developmental series of hybrids
exhibited when three kinds of differentiating characters are conjoined
in them. The hybrids form eight various kinds of egg and pollen
cells--_ABC_, _ABc_, _AbC_, _Abc_, _aBC_, _aBc_, _abC_, _abc_--and each
pollen form unites itself again on the average once with each form of
egg cell.
The law of combination of different characters which governs the
development of the hybrids finds therefore its foundation and
explanation in the principle enunciated, that the hybrids produce egg
cells and pollen cells which in equal numbers represent all constant
forms which result from the combinations of the characters brought
together in fertilisation.
EXPERIMENTS WITH HYBRIDS OF OTHER SPECIES OF PLANTS.
It must be the object of further experiments to ascertain whether
the law of development discovered for _Pisum_ applies also to the
hybrids of other plants. To this end several experiments were recently
commenced. Two minor experiments with species of _Phaseolus_ have been
completed, and may be here mentioned.
An experiment with _Phaseolus vulgaris_ and _Phaseolus nanus_ gave
results in perfect agreement. _Ph. nanus_ had together with the dwarf
axis simply inflated green pods. _Ph. vulgaris_ had, on the other hand,
an axis 10 feet to 12 feet high, and yellow coloured pods, constricted
when ripe. The ratios of the numbers in which the different forms
appeared in the separate generations were the same as with _Pisum_.
Also the development of the constant combinations resulted according to
the law of simple combination of characters, exactly as in the case of
_Pisum_. There were obtained--
Constant Axis Colour of Form of
combinations the unripe pods. the ripe pods.
Public-domain text, read in full here on John Shaqi.
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