Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
With regard to the form of the hybrids and their development, as a rule
an agreement with the observations made in _Pisum_ is unmistakable. It
is otherwise with the exceptional cases cited. Gärtner confesses even
that the exact determination whether a form bears a greater resemblance
to one or to the other of the two original species often involved
great difficulty, so much depending upon the subjective point of view
of the observer. Another circumstance could, however, contribute to
render the results fluctuating and uncertain, despite the most careful
observation and differentiation; for the experiments plants were mostly
used which rank as good species and are differentiated by a large
number of characters. In addition to the sharply defined characters,
where it is a question of greater or less similarity, those characters
must also be taken into account which are often difficult to define
in words, but yet suffice, as every plant specialist knows, to give
the forms a strange appearance. If it be accepted that the development
of hybrids follows the law which is valid for _Pisum_, the series
in each separate experiment must embrace very many forms, since the
number of the components, as is known, increases with the number of
the differentiating characters in _cubic ratio_. With a relatively
small number of experimental-plants the result therefore could only be
approximately right, and in single cases might fluctuate considerably.
If, for instance, the two original stocks differ in seven characters,
and 100 and 200 plants were raised from the seeds of their hybrids to
determine the grade of relationship of the offspring, we can easily see
how uncertain the decision must become, since for seven differentiating
characters the combination series contains 16,384 individuals under
2187 various forms; now one and then another relationship could assert
its predominance, just according as chance presented this or that form
to the observer in a majority of cases.
If, furthermore, there appear among the differentiating characters at
the same time dominant characters, which are transferred entire or
nearly unchanged to the hybrids, then in the terms of the developmental
series that one of the two original stocks which possesses the
majority of dominant characters must always be predominant. In the
experiment described relative to _Pisum_, in which three kinds of
differentiating characters were concerned, all the dominant characters
belonged to the seed parent. Although the terms of the series in their
internal composition approach both original stock plants equally,
in this experiment the type of the seed parent obtained so great
a preponderance that out of each sixty-four plants of the first
generation fifty-four exactly resembled it, or only differed in one
character. It is seen how rash it may be under such circumstances to
draw from the external resemblances of hybrids conclusions as to their
internal nature.
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