Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
Since the form _abc_ occurs once in the series of eight components,
it is consequently little likely that it would be missing among the
experimental plants, even were these raised in a smaller number,
and the transformation would be perfected already by a second
fertilisation. If by chance it did not appear, then the fertilisation
must be repeated with one of those forms nearest akin, _Aabc_, _aBbc_,
_abCc_. It is perceived that such an experiment must extend the farther
_the smaller the number of experimental plants and the larger the
number of differentiating characters_ in the two original species;
and that, furthermore, in the same species there can easily occur a
delay of one or even of two generations such as Gärtner observed.
The transformation of widely divergent species could generally only
be completed in five or six years of experiment, since the number of
different egg cells which are formed in the hybrid increases in square
ratio with the number of differentiating characters.
Gärtner found by repeated experiments that the respective period of
transformation varies in many species, so that frequently a species
_A_ can be transformed into a species _B_ a generation sooner
than can species _B_ into species _A_. He deduces therefrom that
Kölreuter’s opinion can hardly be maintained that “the two natures
in hybrids are perfectly in equilibrium.” It appears, however, that
Kölreuter does not merit this criticism, but that Gärtner rather has
overlooked a material point, to which he himself elsewhere draws
attention, viz. that “it depends which individual is chosen for further
transformation.” Experiments which in this connection were carried out
with two species of _Pisum_ demonstrated that as regards the choice of
the fittest individuals for the purpose of further fertilisation it
may make a great difference which of two species is transformed into
the other. The two experimental plants differed in five characters,
while at the same time those of species _A_ were all dominant and
those of species _B_ all recessive. For mutual transformation _A_
was fertilised with pollen of _B_, and _B_ with pollen of _A_, and
this was repeated with both hybrids the following year. With the
first experiment _B_/_A_ there were eighty-seven plants available in
the third year of experiment for the selections of individuals for
further crossing, and these were of the possible thirty-two forms;
with the second experiment _A_/_B_ seventy-three plants resulted,
which _agreed throughout perfectly in habit with the pollen parent_;
in their internal composition, however, they must have been just as
varied as the forms of the other experiment. A definite selection was
consequently only possible with the first experiment; with the second
some plants selected at random had to be excluded. Of the latter only
a portion of the flowers were crossed with the _A_ pollen, the others
were left to fertilise themselves. Among each five plants which were
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