Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
selected in both experiments for fertilisation there agreed, as the
following year’s culture showed, with the pollen parent:--
1st Experiment. 2nd Experiment.
2 plants -- in all characters
3 " -- " 4 "
-- 2 plants " 3 "
-- 2 " " 2 "
-- 1 plant " 1 character
In the first experiment, therefore, the transformation was completed;
in the second, which was not continued further, two more fertilisations
would probably have been required.
Although the case may not frequently occur that the dominant characters
belong exclusively to one or the other of the original parent plants,
it will always make a difference which of the two possesses the
majority. If the pollen parent shows the majority, then the selection
of forms for further crossing will afford a less degree of security
than in the reverse case, which must imply a delay in the period of
transformation, provided that the experiment is only considered as
completed when a form is arrived at which not only exactly resembles
the pollen plant in form, but also remains as constant in its progeny.
Gärtner, by the results of these transformation experiments, was led to
oppose the opinion of those naturalists who dispute the stability of
plant species and believe in a continuous evolution of vegetation. He
perceives in the complete transformation of one species into another
an indubitable proof that species are fixed within limits beyond which
they cannot change. Although this opinion cannot be unconditionally
accepted we find on the other hand in Gärtner’s experiments a
noteworthy confirmation of that supposition regarding variability of
cultivated plants which has already been expressed.
Among the experimental species there were cultivated plants, such as
_Aquilegia atropurpurea_ and _canadensis_, _Dianthus caryophyllus_,
_chinensis_, and _japonicus_, _Nicotiana rustica_ and _paniculata_, and
hybrids between these species lost none of their stability after four
or five generations[49].
[49] [The argument of these two last paragraphs appears to be that
though the general mutability of natural species might be doubtful,
yet among cultivated plants the transference of characters may be
accomplished, and may occur by integral steps until one species is
definitely “transformed” into the other.]
ON HIERACIUM-HYBRIDS OBTAINED BY ARTIFICIAL FERTILISATION
By G. Mendel.
(_Communicated to the Meeting 9 June, 1869[50]._)
[50] [Published in _Verh. naturf. Ver. Brünn, Abhandlungen_, VIII.
1869, p. 26, which appeared in 1870.]
Although I have already undertaken many experiments in fertilisation
between species of _Hieracium_, I have only succeeded in obtaining the
following 6 hybrids, and only from one to three specimens of them.
Public-domain text, read in full here on John Shaqi.
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