Very careful inquiry would be needed before such a possibility could be negatived. For example, we know that the Laws based on Ancestry can apply to _alternative_ inheritance; witness the case of the Basset-hounds. Here there is no simple Mendelian dominance; but are we sure there is no purity of germ-cells? The new conception goes a long way and it may well reach to such facts as these. But for the present we will assume that Mendel’s principle applies only to _certain phenomena of alternative inheritance_, which is as far as our warrant yet runs. No close student of the recent history of evolutionary thought needs to be told what the attitude of Professor Weldon and his followers has been towards these same disquieting and unwelcome phenomena of alternative inheritance and discontinuity in variation. Holding at first each such fact for suspect, then treating them as rare and negligible occurrences, he and his followers have of late come slowly to accede to the facts of discontinuity a bare and grudging recognition in their scheme of evolution[62]. [62] Read in this connexion Pearson, K., _Grammar of Science_, 2nd ed. 1900, pp. 390–2. Professor Weldon even now opens his essay with the statement--or perhaps reminiscence--that “it is perfectly possible and indeed probable that the difference between these forms of inheritance [blended, mosaic, and alternative] is only one of degree.” This may be true; but reasoning favourable to this proposition could equally be used to prove the difference between mechanical mixture and chemical combination to be a difference of degree. Therefore on the announcement of that discovery which once and for all ratifies and consolidates the conception of discontinuous variation, and goes far to define that of alternative inheritance, giving a finite body to what before was vague and tentative, it is small wonder if Professor Weldon is disposed to criticism rather than to cordiality. * * * * * We have now seen what is the essence of Mendel’s discovery based on a series of experiments of unequalled simplicity which Professor Weldon does not venture to dispute. II. MENDEL AND THE CRITIC’S VERSION OF HIM. _The “Law of Dominance.”_ I proceed to the question of dominance which Professor Weldon treats as a prime issue, almost to the virtual concealment of the great fact of gametic purity. Cross-breds in general, _AB_ and _BA_, named above, may present many appearances. They may all be indistinguishable from _A_, or from _B_; some may appear _A_’s and some _B_’s; they may be patchworks of both; they may be blends presenting one or many grades between the two; and lastly they _may have an appearance special to themselves_ (_being in the latter case, as it often happens, “reversionary”_), a possibility which Professor Weldon does not stop to consider, though it is the clue that may unravel many of the facts which mystify him now.