Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
In the year 1900 Messrs Vilmorin-Andrieux & Co. were kind enough to
send to the Cambridge Botanic Garden on my behalf a set of samples
of the varieties of _Pisum_ and _Phaseolus_, an exhibit of which
had greatly interested me at the Paris Exhibition of that year. In
the past summer I grew a number of these and made some preliminary
cross-fertilizations among them (about 80 being available for these
deductions) with a view to a future study of certain problems,
Mendelian and others. In this work I had the benefit of the assistance
of Miss Killby of Newnham College. Her cultivations and crosses were
made independently of my own, but our results are almost identical. The
experience showed me, what a naturalist would expect and practical men
know already, that _a great deal turns on the variety used_; that some
varieties are very sensitive to conditions while others maintain their
type sturdily; that in using certain varieties Mendel’s experience
as to dominance is regularly fulfilled, while in the case of other
varieties irregularities and even some contradictions occur. That the
dominance of yellow cotyledon-colour over green, and the dominance of
the smooth form over the wrinkled, is a _general_ truth for _Pisum
sativum_ appears at once; that it is a universal truth I cannot believe
any competent naturalist would imagine, still less assert. Mendel
certainly never did. When he speaks of the “law” or “laws” that he
has established for _Pisum_ he is referring to his own discovery of
the purity of the germ-cells, that of the statistical distribution of
characters among them, and the statistical grouping of the different
germ-cells in fertilization, and not to the “Law of Dominance” which he
never drafted and does not propound.
The issue will be clearer if I here state briefly what, as far
as my experience goes, are the facts in regard to the characters
_cotyledon-colour_ and _seed-shapes_ in peas. I have not opportunity
for more than a passing consideration of the _seed-coats_ of pure
forms[63]; that is a maternal character, a fact I am not sure Professor
Weldon fully appreciates. Though that may be incredible, it is evident
from many passages that he has not, in quoting authorities, considered
the consequences of this circumstance.
[63] The whole question as to seed-coat colour is most complex.
Conditions of growth and ripening have a great effect on it. Mr
Arthur Sutton has shown me samples of _Ne Plus Ultra_ grown in
England and abroad. This pea has yellow cotyledons with seed-coats
either yellow or “blue.” The foreign sample contained a much greater
proportion of the former. He told me that generally speaking this is
the case with samples ripened in a hot, dry climate.
Unquestionable Xenia appears occasionally, and will be spoken of
later. Moreover to experiment with such a _plant_-character an extra
generation has to be sown and cultivated. Consequently the evidence
is meagre.
Public-domain text, read in full here on John Shaqi.
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