Mendel's principles of heredity: A defenceMendel, Gregor
Science
Mendel's principles of heredity: A defence
Mendel, Gregor
Heredity; Hybridization; Mendel's law; Weldon, Walter Frank Raphael. Mendel's laws of alternative inheritance in peas
[65] I notice that Vilmorin in the well-known _Plantes Potagères_,
1883, classifies the intermediate-coloured peas with the _green_.
[66] Similarly though _tall_ and _dwarf_ are Mendelian characters,
peas occur of all heights and are usually classified as tall,
half-dwarfs, and dwarfs.
Lastly in some varieties there are many piebalds or mosaics.
From what has been said it will be evident that the description of a
pea in an old book as having been green, blue, white, and so forth,
unless the cotyledon-colour is distinguished from seed-coat colour,
needs careful consideration before inferences are drawn from it.
_Shape._
In regard to shape, if we keep to ordinary shelling peas, the facts are
somewhat similar, but as shape is probably more sensitive to conditions
than cotyledon-colour (not than _seed-coat_ colour) there are
irregularities to be perhaps ascribed to this cause. Broadly, however,
there are two main divisions, round and wrinkled. It is unquestioned
that between these two types every intermediate occurs. Here again
a vast number of varieties can be at once classified into round and
wrinkled (the classification commonly used), others are intermediate
normally. Here also I suspect some fairly clear sub-divisions might be
made in the wrinkled group and in the round group too, but I would not
assert this as a fact.
I cannot ascertain from botanists what is the nature of the difference
between round and wrinkled peas, though no doubt it will be easily
discovered. In maize the round seeds contain much unconverted starch,
while in the wrinkled or sugar-maize this seems to be converted in
great measure as the seed ripens; with the result that, on drying, the
walls collapse. In such seeds we may perhaps suppose that the process
of conversion, which in round seeds takes place on germination, is
begun earlier, and perhaps the variation essentially consists in the
premature appearance of the converting ferment. It would be most rash
to suggest that such a process may be operating in the pea, for the
phenomenon may have many causes; but however that may be, there is
evidently a difference of such a nature that when the water dries out
of the seed on ripening, its walls collapse[67]; and this collapse may
occur in varying degrees.
[67] Wrinkling must of course be distinguished further from the
squaring due to the peas pressing against each other in the pod.
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