Mendelism: Third EditionPunnett, Reginald Crundall
Science
Mendelism: Third Edition
Punnett, Reginald Crundall
Mendel's law
whether we are prepared to recognise facts, and to modify or even destroy
some of the conventions which we have become accustomed to regard as the
foundations of our social life. Whatever be the outcome, there can be
little doubt that the future of our civilisation, perhaps even the
possibility of a future at all, is wrapped up with the recognition we
accord to those who live unseen and inarticulate within us--the fateful
race of gametes so irrevocably bound to us by that closest of all ties,
heredity.
* * * * * {187}
APPENDIX
As some readers may possibly care to repeat Mendel's experiments for
themselves, a few words on the methods used in crossing may not be
superfluous. The flower of the pea with its standard, wings, and median
keel is too familiar to need description. Like most flowers it is
hermaphrodite. Both male and female organs occur on the same flower, and
are covered by the keel. The anthers, ten in number, are arranged in a
circle round the pistil. As soon as they are ripe they burst and shed their
pollen on the style. The pollen tubes then penetrate the stigma, pass down
the style, and eventually reach the ovules in the lower part of the pistil.
Fertilisation occurs here. Each ovule, which is reached by a pollen tube,
swells up and becomes a seed. At the same time the fused carpels enclosing
the ovules enlarge to form the pod. When this, the normal mode of
fertilisation, takes place, the flower is said to be SELFED.
In crossing, it is necessary to emasculate a flower on the plant chosen to
be the female parent. For this purpose a young flower must be taken in
which the anthers have not yet burst. The keel is depressed, and the
stamens bearing the anthers are removed at their base by a {188} pair of
fine forceps. It will probably be found necessary to tear the keel slightly
in order to do this. The pistil is then covered up again with the keel, and
the flower is enclosed in a bag of waxed paper until the following day. The
stigma is then again exposed and dusted with ripe pollen from a flower of
the plant selected as the male parent. This done, the keel is replaced, and
the flower again enclosed in its bag to protect it from the possible
attentions of insects until it has set seed. The bag may be removed in
about a week after fertilisation. It is perhaps hardly necessary to add
that strict biological cleanliness must be exercised during the fertilising
operations. This is readily attained by sterilising fingers and forceps
with a little strong spirit before each operation, thereby ensuring the
death of any foreign pollen grains which may be present.
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