Mendelism: Third EditionPunnett, Reginald Crundall
Science
Mendelism: Third Edition
Punnett, Reginald Crundall
Mendel's law
For this reason the biologist has been
accustomed to demand a very large supply of time, often a great deal more
than the physicist is {150} disposed to grant, and this has sometimes led
him to expostulate with the latter for cutting off the supply. On the newer
views, however, this difficulty need not arise, for we realise that the
origin and establishing of a new form may be a very much more rapid process
than has hitherto been deemed possible.
One last question with regard to evolution. How far does Mendelism help us
in connection with the problem of the origin of species? Among the plants
and animals with which we have dealt we have been able to show that
distinct differences, often considerable, in colour, size, and structure,
may be interpreted in terms of Mendelian factors. It is not unlikely that
most of the various characters which the systematist uses to mark off one
species from another, the so-called specific characters, are of this
nature. They serve as convenient labels, but are not essential to the
conception of species. A systematist who defined the wild sweet pea could
hardly fail to include in his definition such characters as the procumbent
habit, the tendrils, the form of the pollen, the shape of the flower, and
its purple colour. Yet all these and other characters have been proved to
depend upon the presence of definite factors which can be removed by
appropriate crossing. By this means we can produce a small plant a few
inches in height with an erect habit of growth, without tendrils, with
round instead of oblong pollen, and with colourless deformed flowers quite
different {151} in appearance from those of the wild form. Such a plant
would breed perfectly true, and a botanist to whom it was presented, if
ignorant of its origin, might easily relegate it to a different genus.
Nevertheless, though so widely divergent in structure, such a plant must
yet be regarded as belonging to the species _Lathyrus odoratus_. For it
still remains fertile with the many different varieties of sweet pea. It is
not visible attributes that constitute the essential difference between one
species and another. The essential difference, whatever it may be, is that
underlying the phenomenon of sterility. The visible attributes are those
made use of by the systematist in cataloguing the different forms of animal
and plant life, for he has no other choice. But it must not be forgotten
that they are often misleading. Until they were bred together _Euralia
wahlbergi_ and _E. mima_ were regarded as perfectly valid species, and
there is little doubt that numbers of recognised species will eventually
fall to the ground in the same way as soon as we are in a position to apply
the test of breeding. Mendelism has helped us to realise that specific
characters may be but incidental to a species--that the true criterion of
what constitutes a species is sterility, and that particular form of
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