and whites only, while others, again, and these the least numerous, will give nothing but picotees. The new variety is already fixed in a certain definite proportion of the plants; in this particular instance in 1 out of every 27. All that remains to be done is to pick out these plants. Since all the picotees look alike, whatever their breeding capacity, the only way to do this is to save the seed from a number of such plants _individually_, and to raise a further generation. Some of them will be found to breed true. The variety is then established, and may at once be put on the market with full confidence that it will hereafter throw none of the other forms. The all-important thing is to save and sow the seed of separate individuals separately. However alike they look, the seed from different individuals must on no account be mixed. Provided that due care is taken in this respect no long and tedious process of selection is required for the fixation of any given variety. Every possible variety arising from a cross appears in the F_2 generation if only a sufficient {155} number is raised, and of all these different varieties a certain proportion of each is already fixed. Heredity is a question of individuals, and the recognition of this will save the breeder much labour, and enable him to fix his varieties in the shortest possible time. Such cases as these of the sweet pea throw a fresh light upon another of the breeder's conceptions, that of purity of type. Hitherto the criterion of a "pure-bred" thing, whether plant or animal, has been its pedigree, and the individual was regarded as more or less pure bred for a given quality according as it could show a longer or shorter list of ancestors possessing this quality. To-day we realise that this is not essential. The pure-bred picotee appears in our F_2 family though its parent was a purple bicolor, and its remoter ancestors whites for generations. So also from the cross between pure strains of black and albino rabbits we may obtain in the F_2 generation animals of the wild agouti colour which breed as true to type as the pure wild rabbit of irreproachable pedigree. The true test of the pure breeding thing lies not in its ancestry but in the nature of the gametes which have gone to its making. Whenever two similarly constituted gametes unite, whatever the nature of the parents from which they arose, the resulting individual is homozygous in all respects and must consequently breed true. In deciding questions of purity it is to the gamete, and not to ancestry, that our appeal must henceforth be made. {156}