These different species occur round Victoria Nyanza and also on some of the
islands in the lake. Some {127} interesting points are brought out by a
comparison of the occurrence and variation of the species on the mainland
with what is found on Bugalla Island in the Sesse Archipelago. On the
mainland the Pseudacraeas are abundant but the Planemas even more so,
outnumbering the former by about 5:2[83]. Moreover, it is rare to find
individuals more or less intermediate between the three forms, though they
are known to occur. On Bugalla Island, however, a different state of things
is found. The Pseudacraeas are very abundant, whereas the Planemas, owing
doubtless to the scarcity of their food plant, are relatively rare, and are
very greatly outnumbered by the Pseudacraeas. At the same time the
proportion of transitional forms among the Pseudacraeas is definitely
higher than on the mainland. These facts are interpreted by Carpenter as
follows:--
On the mainland where the models are abundant there is a vigorous action on
the part of natural selection. The mimetic forms have a strong advantage
and the non-mimetics have been gradually weeded out. But on the island,
where the Pseudacraeas outnumber the models, the advantage obtained through
mimicry is not so great. The so-called transitional forms are little, if at
all, worse off than those closely resembling the scarce models, and
consequently have as good a chance of surviving as any of the typical
mimetic forms. On {128} the mainland, however, the enemies of _Pseudacraea_
are well acquainted with the _Planema_ models which are here common, and
discriminate against individuals which are not close mimics of the
Planemas. The result is that on the mainland transitional forms are scarcer
than on the island. Natural selection maintains a high standard for the
mimetic likeness on the mainland owing to the abundance of the model; but
when the model is scarce the likeness ceases to be kept up to the mark
strictly, and tends to become lost owing to the appearance of fresh
variations which are no longer weeded out.
Public-domain text, read in full here on John Shaqi.
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