likeness is the distribution of the black pigment with reference to the
lighter pigments; and although the latter are chemically distinct {151} in
the Pierids as compared with other butterflies, there would seem to be no
reason why the same factors governing the distribution of black should not
be common to members of different groups. A distribution of black pigment
similar to that found in a model and its mimic may occur also in a
non-mimetic ally of the mimic. _Dismorphia astynome_, for example,
resembles the Ithomiine _Mechanitis lysimnia_ (Pl. XV, fig. 8) both in the
distribution of black as well as of yellow and bright brown pigments. A
similar distribution of the black pigment is also found in _Dismorphia
avonia_, but the yellow and bright brown of the other two species is here
replaced with white. By a slight though definite alteration in chemical
composition this white pigment could be changed into bright brown and
yellow with the result that _D. avonia_ would closely resemble _D.
astynome_ in its colour scheme and would in this way also become a mimic of
_Mechanitis lysimnia_. Another good instance is that of the females of
_Perrhybris demophile_ and _P. lorena_, the former being black and white,
whereas in the latter the white is replaced by yellow and bright brown,
giving the insect a typical Ithomiine appearance[92]. Here again a definite
small change in the composition of the pigment laid down in the scales
would result in the establishing of a mimetic likeness where there would
otherwise be not even a suggestion of it. It is in accordance with what we
know to-day of variation {152} that such a change should appear suddenly,
complete from the start. And if so there is no difficulty in supposing that
it might be of some advantage to its possessor through the resemblance to
an unpalatable form. Even were the advantage but a slight one it is clear
from previous discussion (p. 96) that the new variety would more or less
rapidly replace the form from which it had sprung. With the continued
operation of natural selection the new form would entirely supplant the
original one, but it is not impossible that in some cases the selecting
agent may be removed before this result has been achieved. In this event
the proportions of the new and the old form would fall into a condition of
equilibrium as in _P. polytes_ in Ceylon, until some other selective agent
arose to disturb the balance. On this view natural selection is a real
factor in connection with mimicry, but its function is to conserve and
render preponderant an already existing likeness, not to build up that
likeness through the accumulation of small variations, as is so generally
assumed. Recent researches in heredity and variation all point to this
restriction of the scope of natural selection. Hitherto an argument in
favour of the older view has been that derived from the study of
adaptation--of an apparent purpose, which, at first sight, appears to be
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