P.S.--If you should visit the Epipactis again, would you gather a few of
the lower flowers which have been opened for some time and have begun
to wither a little, and observe whether pollen is well cleared out of
anther-case. I have been struck with surprise that in nearly all the
lower flowers sent by you, though much of the pollen has been removed,
yet a good deal of pollen is left wasted within the anthers. I observed
something of this kind in Cephalanthera grandiflora. But I fear that you
will think me an intolerable bore.
LETTER 596. TO A.G. MORE. Down, August 5th, 1860.
I am infinitely obliged for your most clearly stated observations on
the bee-orchis. It is now perfectly clear that something removes the
pollen-masses far more with you than in this neighbourhood. But I am
utterly puzzled about the foot-stalk being so often cut through. I
should suspect snails. I yesterday found thirty-nine flowers, and of
them only one pollen-mass in three flowers had been removed, and as
these were extremely much-withered flowers I am not quite sure of
the truth of this. The wind again is a new element of doubt. Your
observations will aid me extremely in coming to some conclusion. (596/1.
Mr. More's observations on the percentage of flowers in which the
pollinia were absent are quoted in "Fertilisation of Orchids," Edition
I., page 68.) I hope in a day or two to receive some day-moths, on the
probosces of which I am assured the pollen-masses of the bee-orchis
still adhere (596/2. He was doomed to disappointment. On July 17th,
1861, he wrote to Mr. More:--"I found the other day a lot of bee-Ophrys
with the glands of the pollinia all in their pouches. All facts point
clearly to eternal self-fertilisation in this species; yet I cannot
swallow the bitter pill. Have you looked at any this year?")...
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