I see that Rhexia grows in Massachusetts; and I suppose has two
different sets of stamens. Now, if in your power, would you observe the
position of the pistil in different plants, in lately opened flowers
of the same age? (I specify this because in Monochaetum I find great
changes of position in the pistils and stamens, as flower gets old).
Supposing that my prophecy should turn out right, please observe whether
in both forms the passage into the flower is not [on] the upper side
of the pistil, owing to the basal part of the pistil lying close to the
ring of filaments on the under side of the flower. Also I should like to
know the colour of the two sets of anthers. This would take you only a
few minutes, and is the only way I see that I can find out whether these
plants are dimorphic in this peculiar way--i.e., only in the position
of the pistil (621/1. In Exacum and in Saintpaulia the flowers are
dimorphic in this sense: the style projects to either the right or the
left side of the corolla, from which it follows that a right-handed
flower would fertilise a left-handed one, and vice versa. See Willis,
"Flowering Plants and Ferns," 1897, Volume I., page 73.) and in its
relation to the two kinds of pollen. I am anxious about this, because if
it should prove so, it will show that all plants with longer and shorter
or otherwise different anthers will have to be examined for dimorphism.
LETTER 622. TO ASA GRAY. March 15th [1862].
...I wrote some little time ago about Rhexia; since then I have been
carefully watching and experimenting on another genus, Monochaetum; and
I find that the pistil is first bent rectangularly (as in the sketch
sent), and then in a few days becomes straight: the stamens also move.
If there be not two forms of Rhexia, will you compare the position of
the part in young and old flowers? I have a suspicion (perhaps it will
be proved wrong when the seed-capsules are ripe) that one set of anthers
are adapted to the pistil in early state, and the other set for it
in its later state. If bees visit the Rhexia, for Heaven's sake watch
exactly how the anther and stigma strike them, both in old and young
flowers, and give me a sketch.
Again I say, do not hate me.
LETTER 623. TO J.D. HOOKER. Leith Hill Place, Dorking, Thursday, 15th
[May 1862].
Public-domain text, read in full here on John Shaqi.
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