I am much obliged for your letter, which is full of interesting matter.
I shall be very glad to look at the capsule of the Acropera when
ripe, and pray present my thanks to Mr. MacNab. (634/1. See Letter 608
(Lindley, December 15th, 1861). Also "Fertilisation of Orchids," Edition
II., page 172, for an account of the observations on Acropera which were
corrected by Scott.) I should like to keep it till I could get a capsule
of some other member of the Vandeae for comparison, but ultimately all
the seeds shall be returned, in case you would like to write any
notice on the subject. It was, as I said (634/2. Letter 633.), only
"in desperation" that I suggested that the flower might be a male and
occasionally capable of producing a few seeds. I had forgotten Gartner's
remark; in fact, I know only odds and ends of Botany, and you know far
more. One point makes the above view more probable in Acropera than in
other cases, viz. the presence of rudimentary placentae or testae, for
I cannot hear that these have been observed in the male plants. They do
not occur in male Lychnis dioica, but next spring I will look to male
holly flowers. I fully admit the difficulty of similarity of stigmatic
chamber in the two Acroperas. As far as I remember, the blunt end of
pollen-mass would not easily even stick in the orifice of the chamber.
Your view may be correct about abundance of viscid matter, but seems
rather improbable. Your facts about female flowers occurring where males
alone ought to occur is new to me; if I do not hear that you object, I
will quote the Zea case on your authority in what I am now writing on
the varieties of the maize. (634/3. See "Animals and Plants," Edition
II., Volume I., page 339: "Mr. Scott has lately observed the rarer case
of female flowers on a true male panicle, and likewise hermaphrodite
flowers." Scott's paper on the subject is in "Trans. Bot. Soc.
Edinburgh," Volume VIII. See Letter 151, Volume I.) I am glad to hear
that you are now working on the most curious subject of parthenogenesis.
I formerly fancied that I observed female Lychnis dioica seeded without
pollen. I send by this post a paper on Primula, which may interest you.
(634/4. "Linn. Soc. Journal," 1862.) I am working on the subject, and if
you should ever observe any analogous case I should be glad to hear. I
have added another very clever pamphlet by Prof. Asa Gray. Have you a
copy of my Orchis book? If you have not, and would like one, I should be
pleased to send one. I plainly see that you have the true spirit of an
experimentalist and good observer. Therefore, I ask whether you have
ever made any trials on relative fertility of varieties of plants (like
those I quote from Gartner on the varieties of Verbascum). I much
want information on this head, and on those marvellous cases (as some
Lobelias and Crinum passiflora) in which a plant can be more easily
fertilised by the pollen of another species than by its own good pollen.
I am compelled to write in haste.
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