An extraordinary amount of ingenuity has been expended in this connection,
in order to avoid here, where perhaps it is most difficult of all, the use
of “teleological” principles, and to remain faithful to the orthodox,
exclusively mechanical mode of interpretation. To this category belong
Darwin’s gemmules, Haeckel’s plastidules, Nägeli’s micellæ, Weismann’s
labyrinth of ids, determinants, and biophors within the germ-plasm, and
Roux’s ingenious hypothesis of the struggle of parts, which is an attempt
to apply the Darwinian principle within the organism in order here also to
rebut the teleological interpretation by giving a scientific one.(66)
Heredity.
6. With this fifth line of thought a sixth is associated and intertwined.
The problem of development is closely bound up with that of “heredity.” A
developing organism follows the parental type. The acorn in its growth
follows the type of the parent oak, repeating all its morphological and
physiological characters down to the most intimate detail. And the animal
organism adds to this also the whole psychical equipment, the instincts,
the capacities of will and consciousness which distinguish its parents.
The problems of the fifth and sixth order are closely inter-related, the
sixth problem being in reality the same as the fifth, only in greater
complexity.
A step towards the mechanical solution of this problem was indicated in
the “preformation theory” advanced by Leibnitz, and elaborated by Bonnet.
According to this theory the developing organism is enclosed in the
minutest possible form within the egg, and is thus included in the
parental organism, in miniature indeed, but quite complete. Thus the
problem of the “development of form” or of “heredity” was, so to speak,
ruled out of court; all that was assumed was continuous growth and
self-unfolding.
Opposed to this theory was one of later growth, the theory of epigenesis,
which maintained that the organism developed without preformation from the
still undifferentiated and homogeneous substance of the egg. The
supporters of the first theory considered themselves much more scientific
and exact than those of the second. And not without reason. For the theory
of epigenesis obviously required mysterious formative principles, and
equally mysterious powers of recollection and recapitulation, which
impelled the undifferentiated ovum substance into the final form,
precisely like that of its ancestors. Nor need the preformationists have
greatly feared the reproach, that the parental organism must have been
included within the grand-parental, and so on backwards to the first
parents in Paradise. For this “Chinese box” encapsulement theory only
requires that we should grant the idea of the infinitely little, and that
idea is already an integral part of our thinking.
Public-domain text, read in full here on John Shaqi.
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