Darwin himself did not regard these two theories as opposed to the theory
of selection, but utilised them as subsidiary interpretations. It is
obvious, however, that at bottom they conceal an essentially different
fundamental idea, which, if followed out to its logical consequences,
reduces the “struggle for existence” to at most a wholly indifferent
accessory circumstance. Weismann felt this, and hence his entirely
consistent endeavours to show by great examples, such as the origin of
flowers, the mutual adaptations of flowers and insects, the phenomena of
mimicry, and many other cases, that neither the Lamarckian nor any other
factor in evolution, except only natural, passive selection, suffices as
an interpretation. From the Darwinian standpoint he is absolutely right,
and must needs speak of the “omnipotence of natural selection,” for it
must either be omnipotent, or it must give place to the other two factors,
and retain only the significance we attributed to it in another connection
(p. 157), which amounts to saying none at all. It is obvious enough why
the discussion as to these factors should centre round the question of the
“inheritance of acquired characters,” “acquired” either through the use or
disuse of organs, the exercise or non-exercise of functions, or through
the stimuli of the external world.
The neo-Lamarckian conflict with Darwinism has become more and more acute
in recent times, and the neo-Lamarckians have sometimes passed from
contrasting rival interpretations to excluding the Darwinian factor
altogether. As the particular champion of the neo-Lamarckian view, we must
name Th. Eimer, the recently deceased Tübingen zoologist. His chief work
is in three volumes, entitled “Die Entstehung der Arten auf Grund von
Vererbung erworbener Eigenschaften, nach Gesetzen organischen
Wachsens.”(43) It is a polemic against Weismannism in all details, even to
the theory of “germinal selection.” Eimer follows in the footsteps of St.
Hilaire, and shows what a relatively plastic and sensitive creature the
organism is to the surrounding world, the conditions of nutrition and
other such influences. There is in this connection a particularly
instructive chapter on the physiological and other variations brought
about by external influences which act as “stimuli of the nervous system.”
The whole theory of Lamarck and St. Hilaire transcends—notwithstanding the
protests of Eimer to the contrary—the categories of the mechanical theory
of life, and this chapter does so in particular. The array of facts here
marshalled as to the spontaneous self-adaptation of organisms to their
environment—in relation to colour mainly—forms the most thoroughgoing
refutation of Darwinism that it is possible to imagine. It is shown, too,
by a wealth of examples from osteology, how use (and the necessities of
the case—a consideration which again goes beyond the bounds of mere
Lamarckism) may modify, increase or diminish vertebræ, ribs, skull and
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