North American Recent Soft-Shelled Turtles (Family Trionychidae)Webb, Robert G. (Robert Gravem)
Science
North American Recent Soft-Shelled Turtles (Family Trionychidae)
Webb, Robert G. (Robert Gravem)
Soft-shelled turtles
_T. muticus_ is the smallest species in North America; the maximum
size of the plastron in adult males is approximately 14.0 centimeters
(16.0 cm. in _spinifer_) and of adult females 21.5 centimeters (31.0
cm. in _spinifer_). Males and females of _muticus_ are sexually mature
at approximately the same size as some _T. s. emoryi_; also, the great
development of the plastral callosities in _muticus_ corresponds to
that in some _emoryi_. The head is narrower in _muticus_ than in
_ferox_ or _spinifer_. The carapaces of specimens of _muticus_
exceeding plastral lengths of 8.0 centimeters are wider than those of
_ferox_, _ater_, _T. s. emoryi_ and _T. s. guadalupensis_ of
corresponding size. _T. muticus_ differs from _ater_ and three
subspecies of _spinifer_ (_pallidus_, _guadalupensis_, _emoryi_) in
having the carapace widest at a plane approximately one-half the
length of the carapace. The snout is longer in _muticus_ than in
_ferox_ and _spinifer_. _T. muticus_ differs from _ferox_ but
resembles _spinifer_ in having a relatively short plastron.
The skulls of _muticus_ differ from those of _ferox_ but resemble
those of _spinifer_ in usually having the skull widest at the level of
the squamosals. Skulls of _muticus_ resemble those of _ferox_ but
differ from those of _spinifer_ in usually lacking a well-developed
opisthotic-exoccipital spur. Skulls of _muticus_ are different from
those of _ferox_ and _spinifer_ in having the 1) ventral surface of
the supraoccipital spine widest proximally, lacking a medial ridge, 2)
foramen magnum ovoid, 3) distal part of opisthotic wing truncate, 4)
lateral condyle of articular surface of quadrate tapered posteriorly,
smaller than medial articular surface, and 5) maxillaries not in
contact above premaxillaries.
Plastrons of _muticus_ differ from those of _spinifer_ and _ferox_ in
having an obtusely-angled epiplastron, relatively large callosities
in adults, and a wide hyo-hypoplastral bridge (in relation to length).
_Remarks._--Agassiz (1857:399) regarded Lesueur's _Trionyx muticus_ as
the type species of the genus _Amyda_ and the only species known to
belong to the genus _Amyda_. Stejneger (1944:7, 9, 12) proposed the
generic name _Euamyda_ as a new name for the North American _Amyda
mutica_ as understood by Agassiz. _Euamyda_ was proposed for use only
if Agassiz's understanding was found to be correct. Actually,
Stejneger thought that the Old World and New World kinds concerned
were congeneric, and that the type species of the genus _Amyda_ was
the Old World species _Amyda javanica_ Schweigger (= _Testudo
cartilaginea_ Boddaert).
If _Trionyx muticus_ Lesueur is considered to be generically distinct
from other soft-shelled turtles, _Euamyda_ Stejneger, 1944, is
available as a generic name with _Trionyx muticus_ Lesueur, 1827, as
the type species (by monotypy).
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