North American Recent Soft-Shelled Turtles (Family Trionychidae)Webb, Robert G. (Robert Gravem)
Science
North American Recent Soft-Shelled Turtles (Family Trionychidae)
Webb, Robert G. (Robert Gravem)
Soft-shelled turtles
The occurrence of the two clines, _spinifer-hartwegi_ and
_pallidus-guadalupensis_, in the species _spinifer_ are notable in that
the former occurs mostly in one large continuous drainage system, that
of the Mississippi, and shows no sharp break in the one character
distinguishing the two subspecies whereas populations composing the
_pallidus-guadalupensis_ cline are separated into several river
drainages, and show a relatively sharp break in several characters at
the Brazos-Colorado river divide. This situation seemingly supports the
thesis that clines are maintained by some sort of parallel gradient in
ecological or geological conditions. It is notable that streams draining
the Edward's Plateau (inhabited by _guadalupensis_) differ in quantity
(more) and quality (especially CO_{3}^{--}, Ca^{++}, and Mg^{++} ions)
of their solutes, and probably pH (higher) from those farther east
(Hubbs, 1957:102). The gross difference in habitats mentioned above
(sandy, turbid, sluggish streams in the west _vs._ clear, swift streams
in the east) may affect the differentiation recognizable in the
_spinifer-hartwegi_ cline.
Daily and Seasonal Activity
_Diurnal Habits_
Softshells bask on débris in the water or on banks close to the water;
basking presumably raises the bodily temperature. In general in the
southeastern and southwestern United States, I have seen softshells
basking only rarely but once saw six at one time close together on logs
in Bowie Creek, Hattiesburg, Mississippi (species undetermined). Surface
(1908:122) saw _spinifer_ in rows on rocks or logs in tributaries of the
Ohio River. Duellman and Schwartz (1958:271-72) stated that _ferox_
basks on banks or beds of aquatic vegetation. Deckert (1918:31)
mentioned large _ferox_ "sunning in shallow water at edge of pond."
Minton (1944:447) wrote that _muticus_ and _spinifer_ sun on steep mud
banks (Wabash River). Cahn (1937:180) stated that _muticus_ (in
Illinois) basks on banks at the water's edge but seldom on logs, and
suggests that _muticus_ is less prone to leave the water than
_spinifer_. According to Carr (1952:438), _muticus_ never basks on
logs or rocks. In Ohio, Conant (1951:159) mentioned _spinifer_ as
occasionally basking upon a log or rock, or sometimes on steep clay
banks of streams. On banks, quick escape is facilitated by directing the
head toward the water, thus eliminating the time that it would take to
turn around on land (Conant, _loc. cit._; Newman, 1906:129). Evermann
and Clark (1920:593) mentioned _spinifer_ as basking on sandy or grassy
shores, and large boulders. Muller (1921:181) wrote that _muticus_ basks
four to ten feet from the water's edge on gently sloping sand and mud
shores of small islands in the Mississippi River (near Fairport, Iowa).
Muller stated that basking usually occurs in the morning, up until 2 p.
m., and that beaches with a northern exposure were preferred; he
observed 37 turtles within a 50-foot stretch of beach. In captivity,
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