North American Recent Soft-Shelled Turtles (Family Trionychidae)Webb, Robert G. (Robert Gravem)
Science
North American Recent Soft-Shelled Turtles (Family Trionychidae)
Webb, Robert G. (Robert Gravem)
Soft-shelled turtles
I have not studied in detail the many fossil remains but such
examination as I have made of them suggests that many of the characters
used as a basis for distinguishing fossil forms in North America are
subject to individual variation or are of no diagnostic value in the
living species (Hummel, 1929:769). Knowledge of the variation in the
living species of the Old World would aid in adequately appraising the
North American fossils. Some osteological characters of the three living
American species (excluding _ater_) together with data on variation
within a given species are mentioned below. Some differences in skulls
of the three species already were mentioned in the section "Osteological
Characters." Because most fossil remains are those of the carapace and
plastron, attention is here given to those structures.
_Widened alveolar surfaces of jaws._--An ontogenetic variation
affecting large skulls of _T. ferox_ and some individuals of _T.
spinifer asper_; presumably confined to females. Of especial interest
is its presence in some populations of _asper_ that are not otherwise
distinguishable (external characters) from the rest of the individuals
comprising that subspecies.
_Sculpturing._--No differences in pattern (generally of anastomosing
ridges) on carapace or plastron; fineness or coarseness seemingly
correlated with size; frequency and kind (knoblike or ridgelike) of
bony prominences on carapace variable; bony prominences confined to
species _spinifer_ and _ferox_, occurring principally on large
females.
_Fontanelles of carapace._--Closure more or less correlated with
increasing size, although much variation noted between individuals of
same size; small individuals have fontanelles confluent (medially),
thus separating nuchal from contact with first neural and first pair
of pleurals.
_Number and arrangement of neurals and pleurals._--Neurals number six
to nine, usually seven or eight; pleurals number seven or eight pairs,
and may or may not be in contact with each other posteriorly; eighth
pair of pleurals when present reduced, never contacting seventh
neural; arrangement posteriorly variable (see Fig. 16 and Tab. 5).
_Plastral callosities._--Increase in size with advancing age causing
corresponding reduction in size of plastral vacuity; relatively best
developed in _muticus_ (all elements touching medially on KU 41380
leaving no plastral vacuity); probably no callosities on preplastra or
epiplastron of _ferox_; callosity on epiplastron of _spinifer_ not
covering entire surface (as it may in _muticus_).
_Epiplastron._--Obtusely-angled (greater than 90 degrees) in
_muticus_; acutely-angled (90 degrees or less) in _ferox_ and
_spinifer_.
_Hyo-hypoplastral suture._--Usually present, but occasionally absent,
in all species.
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