North American Recent Soft-Shelled Turtles (Family Trionychidae)Webb, Robert G. (Robert Gravem)
Science
North American Recent Soft-Shelled Turtles (Family Trionychidae)
Webb, Robert G. (Robert Gravem)
Soft-shelled turtles
_Comparisons._--_T. s. spinifer_ can be distinguished from all other
subspecies of _T. spinifer_ by the presence of large black ocelli
(diameter 9-10 mm. on adult males, 2-3 mm. on hatchlings) in
combination with only one dark marginal line. _T. s. spinifer_
resembles _asper_ in having ocelli or dots on the carapace but differs
from _asper_ in having only one dark marginal line and larger ocelli.
_T. s. spinifer_ differs from _hartwegi_ only in the large size of the
ocelli. _T. s. spinifer_ resembles _hartwegi_ and _asper_ but differs
from _pallidus_, _guadalupensis_ and _emoryi_ in having blackish spots
and ocelli on the carapace and lacking whitish dots. _T. s. spinifer_
resembles _hartwegi_, _asper_, and _pallidus_ and differs from
_guadalupensis_ and _emoryi_ in having conical or knoblike tubercles
on the anterior edge of the carapace on large females.
_T. s. spinifer_ differs from the subspecies _asper_, _guadalupensis_
and _emoryi_ in having a relatively narrower head, and from _emoryi_
in having a relatively wider carapace. _T. s. spinifer_ resembles
_hartwegi_ and _asper_ but differs from the other subspecies in having
the carapace widest at a plane approximately one-half way back on the
carapace. The subspecies _spinifer_ and _hartwegi_ have longer snouts
than _pallidus_, _guadalupensis_, and _emoryi_. _T. s. spinifer_
differs from _asper_ but resembles all the other subspecies in having
a relatively longer plastron.
_Remarks._--Lesueur's description of _Trionyx spiniferus_
(1827:258-261, Pl. 6) seems to be based mostly, if not entirely, on a
large female (length of carapace, 13 inches), which was "Le plus grand
des individus observes ..." (_op. cit._:258); an accompanying
illustration depicting the dorsal surface of the bony carapace is
unusual in lacking neurals (Pl. 6, E). Duméril and Bibron (1835:481)
mentioned eight or nine additional specimens that Lesueur sent to the
Museum of Natural History in Paris. Dr. Jean Guibé informed me under
letter dated September 24, 1959, that a larger stuffed female, bearing
catalog number 8808 is regarded as the holotype, and that there are
seven additional specimens (1949, 4143, 8807, 8809-12) in the museum
at Paris. All turtles were obtained by Lesueur from the Wabash River.
To my knowledge no specimen that was available to Lesueur has been
specifically designated as a type. Because the description seems to be
based on one specimen, undoubtedly No. 8808, this specimen has been
regarded as the holotype. However, Lesueur referred to several
specimens and did not mention a type in the original description;
consequently I prefer to regard No. 8808 as a lectotype.
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