North American Recent Soft-Shelled Turtles (Family Trionychidae)Webb, Robert G. (Robert Gravem)
Science
North American Recent Soft-Shelled Turtles (Family Trionychidae)
Webb, Robert G. (Robert Gravem)
Soft-shelled turtles
_Comparisons._--_T. s. hartwegi_ can be distinguished from all other
subspecies of _T. spinifer_ by the presence of small dots and ocelli
on the carapace that are all of approximately the same size in
combination with only one dark marginal line. _T. s. hartwegi_
resembles _asper_ in having small blackish ocelli or dots on the
carapace but differs from _asper_ in having only one dark marginal
line. _T. s. hartwegi_ differs from _spinifer_ only in the small size
of the ocelli. _T. s. hartwegi_ resembles _spinifer_ and _asper_, but
differs from _pallidus_, _guadalupensis_ and _emoryi_ in having
blackish spots and ocelli on the carapace and lacking small whitish
spots. _T. s. hartwegi_ resembles _spinifer_, _asper_ and _pallidus_
but differs from _guadalupensis_ and _emoryi_ in having conical or
knoblike tubercles on the anterior edge of the carapace on large
females.
_T. s. hartwegi_ differs from the subspecies _asper_, _guadalupensis_
and _emoryi_ in having a narrower head, and from _emoryi_ in having a
wider carapace. _T. s. hartwegi_ resembles _spinifer_ and _asper_ but
differs from the other subspecies in having the carapace widest at a
plane approximately one-half way back on the carapace. _T. s.
hartwegi_ and _spinifer_ have longer snouts than do _pallidus_ and
_guadalupensis_ or _emoryi_. _T. s. hartwegi_ differs from _asper_ but
resembles the other subspecies in having a relatively longer plastron.
_Remarks._--The validity of _T. s. hartwegi_ has never been
questioned. It intergrades with _spinifer_ over a broad area
paralleling the Mississippi River. For convenience, specimens
occurring west of the Mississippi River are referred to the subspecies
_hartwegi_. Figure 8 shows much variation in size of ocelli on
different individuals from the same state. For example, UMMZ 92667,
plastral length 6.7 centimeters has a uniform pale brown carapace
lacking any dark marks, whereas UMMZ 92652, plastral length 5.9
centimeters has some ocelli three millimeters in diameter on the
carapace. Both are from Iowa. One specimen from Kansas, KU 1954
(Doniphan County, plastral length 11.8 cm.), has ocelli four
millimeters in diameter, and USNM 7648 captured farther west at Fort
Laramie, Wyoming, an adult male having a plastral length of 11.0
centimeters, has some ocelli five millimeters in diameter on the
carapace. TTC 1090, an adult male from the panhandle of Texas has some
ocelli so much as 5.5 millimeters in diameter. The size of the ocelli
seemingly varies in the same local population.
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