On Germinal Selection as a Source of Definite VariationWeismann, August
Science
On Germinal Selection as a Source of Definite Variation
Weismann, August
Variation (Biology)
[11] On the same day on which the present address was delivered at the
International Congress of Zooelogists in Leyden, and on the same occasion,
Dr. W. B. Scott, Professor of Geology in Princeton College, New Jersey,
read a very interesting paper on the tertiary mammalian fauna of North
America, in which, without a knowledge of my paper, he took his stand
precisely on this argument and arrived at the opinion that it could not
possibly be the ordinary individual variations which accomplished phyletic
evolution, but that it was necessary to assume in addition phyletic
variations. I believe our views are not as widely remote as might be
supposed. Of course, I see no reason for assuming two kinds of hereditary
variations, different _in origin_. Still it is likely that only a
relatively small portion of the numberless individual variations lie on the
path of phyletic advancement and so under the _guidance_ of germinal
selection mark out the way of further development; and hence it would be
quite possible in this sense to distinguish continuous, _definitely
directed_ individual variations from such as fluctuate hither and thither
with no uniformity in the course of generations. The root of the two is of
course the same, and they admit of being distinguished from each other only
by their success, phyletic modification, or by their failure.
[12] H. F. Osborn, "The Hereditary Mechanism and the Search for the Unknown
Factors of Evolution," in _Biological Lectures delivered at the Marine
Biolog. Lab. at Wood's Holl in the Summer Session of 1894_. Boston, 1895.
[13] In 1886. See my paper on "Retrogression in Nature," published in
English in Nos. 105, 107, 108, and 109 of _The Open Court_, and also in my
essays on _Heredity_, Jena, 1892.
[14] _Neue Gedanken zur Vererbungsfrage_, Jena, 1895.
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