When the lines of force stream inwards from the periphery towards
a point in the interior of the cell, the particles susceptible of
attraction either crowd towards the surface of the cell, or, when
retarded by friction, are seen forming lines or “fibrillae” which
radiate outwards from the centre and constitute a so-called “aster.” In
the cells of columnar or ciliated epithelium, where the sides of the
cell are symmetrically disposed to their neighbours but the free and
attached surfaces are very diverse from one another in their external
relations, it is these latter surfaces which constitute the opposite
poles; and in accordance with the parallel lines of force so set up,
we very frequently see parallel lines of granules which have ranged
themselves perpendicularly to the free surface of the cell (cf. fig.
97).
A simple manifestation of “polarity” may be well illustrated by the
phenomenon of diffusion, where we may conceive, and may automatically
reproduce, a “field of force,” with its poles and visible lines of
equipotential, very much as in Faraday’s conception of the field of
force of a magnetic system. Thus, in one of Leduc’s experiments[224],
if we spread a layer of salt solution over a level {168} plate of
glass, and let fall into the middle of it a drop of indian ink, or
of blood, we shall find the coloured particles travelling outwards
from the central “pole of concentration” along the lines of diffusive
force, and so mapping out for us a “monopolar field” of diffusion: and
if we set two such drops side by side, their lines of diffusion will
oppose, and repel, one another. Or, instead of the uniform layer of
salt solution, we may place at a little distance from one another a
grain of salt and a drop of blood, representing two opposite poles:
and so obtain a picture of a “bipolar field” of diffusion. In either
case, we obtain results closely analogous to the “morphological,”
but really _dynamical_, polarity of the organic cell. But in all
probability, the dynamical polarity, or asymmetry of the cell is a very
complicated phenomenon: for the obvious reason that, in any system,
one asymmetry will tend to beget another. A chemical asymmetry will
induce an inequality of surface-tension, which will lead directly to a
modification of form; the chemical asymmetry may in turn be due to a
process of electrolysis in a polarised electrical field; and again the
chemical heterogeneity may be intensified into a chemical “polarity,”
by the tendency of certain substances to seek a locus of greater or
less surface-energy. We need not attempt to grapple with a subject
so complicated, and leading to so many problems which lie beyond the
sphere of interest of the morphologist. But yet the morphologist, in
his study of the cell, cannot quite evade these important issues; and
we shall return to them again when we have dealt somewhat with the form
of the cell, and have taken account of some of the simpler phenomena of
surface-tension.
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