connected with the organism as such. They include such manifestations
of the physical forces, in their various permutations and combinations,
as may also be witnessed, under appropriate conditions, in non-living
things.
When we attempt to separate our purely morphological or “purely
embryological” studies from physiological and physical investigations,
we tend _ipso facto_ to regard each particular structure and
configuration as an attribute, or a particular “character,” of this or
that particular organism. From this assumption we are apt to go on to
the drawing of new conclusions or the framing of new theories as to the
ancestral history, the classificatory position, the natural affinities
of the several organisms: in fact, to apply our embryological knowledge
mainly, and at times exclusively, to the study of _phylogeny_.
When we find, as we are not long of finding, that our phylogenetic
hypotheses, as drawn from embryology, become complex and unwieldy, we
are nevertheless reluctant to admit that the whole method, with its
fundamental postulates, is at fault. And yet nothing short of this
would seem to be the case, in regard to the earlier phases at least
of embryonic development. All the evidence at hand goes, as it seems
to me, to shew that embryological data, prior to and even long after
the epoch of segmentation, are essentially a subject for physiological
and physical investigation and have but the very slightest link with
the problems of systematic or zoological classification. Comparative
embryology has its own facts to classify, and its own methods and
principles of classification. Thus we may classify eggs according to
the presence or absence, the paucity or abundance, of their associated
food-yolk, the chromosomes according to their form and their number,
the segmentation according to its various “types,” radial, bilateral,
spiral, and so forth. But we have little right to expect, and in
point of fact we shall very seldom and (as it were) only accidentally
find, that these embryological categories coincide with the lines of
“natural” or “phylogenetic” classification which have been arrived at
by the systematic zoologist.
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Public-domain text, read in full here on John Shaqi.
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