It is far from correct to say, as is often done, that the cell-wall,
or cell-membrane, belongs “to the passive products of protoplasm
rather than to the living cell itself”; or to say that in the animal
cell, the cell-wall, because it is “slightly developed,” is relatively
unimportant compared with the important role which it assumes in
plants. On the contrary, it is quite certain that, whether visibly
differentiated into a semi-permeable membrane, or merely constituted by
a liquid film, the surface of the cell is the seat of {199} important
forces, capillary and electrical, which play an essential part in
the dynamics of the cell. Even in the thickened, largely solidified
cellulose wall of the plant-cell, apart from the mechanical resistances
which it affords, the osmotic forces developed in connection with it
are of essential importance.
But if the cell acts, after this fashion, as a whole, each part
interacting of necessity with the rest, the same is certainly true of
the entire multicellular organism: as Schwann said of old, in very
precise and adequate words, “the whole organism subsists only by means
of the _reciprocal action_ of the single elementary parts[256].”
As Wilson says again, “the physiological autonomy of the individual
cell falls into the background ... and the apparently composite
character which the multicellular organism may exhibit is owing to
a secondary distribution of its energies among local centres of
action[257].”
It is here that the homology breaks down which is so often drawn, and
overdrawn, between the unicellular organism and the individual cell of
the metazoon[258].
Whitman, Adam Sedgwick[259], and others have lost no opportunity of
warning us against a too literal acceptation of the cell-theory,
against the view that the multicellular organism is a colony (or,
as Haeckel called it (in the case of the plant), a “republic”) of
independent units of life[260]. As Goethe said long ago, “Das
lebendige ist zwar in Elemente {200} zerlegt, aber man kann es aus
diesen nicht wieder zusammenstellen und beleben;” the dictum of the
_Cellularpathologie_ being just the opposite, “Jedes Thier erscheint
als eine Summe vitaler Einheiten, von denen _jede den vollen Charakter
des Lebens an sich trägt_.”
Public-domain text, read in full here on John Shaqi.
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