find that a plasmodium of Aethalium, for instance, creeps towards a
damp spot, or towards a warm spot, or towards substances that happen
to be nutritious, and again creeps away from solutions of sugar or of
salt, we seem to be dealing with phenomena every one of which can be
paralleled by ordinary phenomena of surface tension[279]. Even the
soap-bubble itself is imperfectly in equilibrium, for the reason that
its film, like the protoplasm of amoeba or Aethalium, is an excessively
heterogeneous substance. Its surface tensions vary from point to
point, and chemical changes and changes of temperature increase and
magnify the variation. The whole surface of the bubble is in constant
movement as the concentrated portions of the soapy fluid make their way
outwards from the deeper layers; it thins and it thickens, its colours
change, currents are set up in it, and little bubbles glide over it; it
continues in this state of constant movement, as its parts strive one
with another in all their interactions towards equilibrium[280].
In the case of the naked protoplasmic cell, as the amoeboid phase
is emphatically a phase of freedom and activity, of chemical and
physiological change, so, on the other hand, is the spherical form
indicative of a phase of rest or comparative inactivity. In the one
phase we see unequal surface tensions manifested in the creeping
movements of the amoeboid body, in the rounding off of the ends of the
pseudopodia, in the flowing out of its substance over a particle of
“food,” and in the current-motions in the interior of its mass; till
finally, in the other phase, when internal homogeneity and equilibrium
have been attained and the potential {213} energy of the system is for
the time being at a minimum, the cell assumes a rounded or spherical
form, passing into a state of “rest,” and (for a reason which we shall
presently see) becoming at the same time “encysted.”
[Illustration: Fig. 60.]
In a budding yeast-cell (Fig. 60), we see a more definite and
restricted change of surface tension. When a “bud” appears, whether
with or without actual growth by osmosis or otherwise of the mass,
it does so because at a certain part of the cell-surface the surface
tension has more or less suddenly diminished, and the area of that
portion expands accordingly; but in turn the surface tension of the
expanded area will make itself felt, and the bud will be rounded off
into a more or less spherical form.
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