It is obvious that there are innumerable solitary plant-cells, and
unicellular organisms in general, which, like the yeast-cell, do not
correspond to any of the simple forms that may be generated under the
influence of simple and homogeneous surface-tension; and in many cases
these forms, which we should expect to be unstable and transitory,
have become fixed and stable by reason of the comparatively sudden or
rapid solidification of the envelope. This is the case, for instance,
in many of the more complicated forms of diatoms or of desmids, where
we are dealing, in a less striking but even more curious way than in
the budding yeast-cell, not with one simple act of formation, but
with a complicated result of successive stages of localised growth,
interrupted by phases of partial consolidation. The original cell has
acquired or assumed a certain form, and then, under altering conditions
and new distributions of energy, has thickened here or weakened there,
and has grown out or tended (as it were) to branch, at particular
points. We can often, or indeed generally, trace in each particular
stage of growth or at each particular temporary growing point, the laws
of surface tension manifesting themselves in what is for the time being
a fluid surface; nay more, even in the adult and completed structure,
we have little difficulty in tracing and recognising (for instance
in the outline of such a desmid as Euastrum) the rounded lobes that
have successively grown or flowed out from the original rounded and
flattened cell. What we see in a many chambered foraminifer, such as
Globigerina or Rotalia, is just the same thing, save that it is carried
out in greater completeness and perfection. The little organism as a
whole is not a figure of equilibrium or of minimal area; but each new
bud or separate chamber is such a figure, conditioned by the forces of
surface tension, and superposed upon the complex aggregate of similar
bubbles after these latter have become consolidated one by one into a
rigid system.
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Public-domain text, read in full here on John Shaqi.
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