But this phenomenon, while it brings about a certain departure from
complete symmetry, is still compatible with, and coexistent with,
many of the phenomena which we have seen to be associated with
surface-tension. The symmetry of tensions still leaves the cell a solid
of revolution, and its surface is still a surface of equilibrium. The
fluid pressure within the cylinder still causes the film or membrane
which caps its ends to be of a spherical form. And in the young cell,
where the surface pellicle is absent or but little differentiated, as
for instance in the oögonium of Achlya, or in the young zygospore of
Spirogyra, we always see the tendency of the entire structure towards
a spherical form reasserting itself: unless, as in the latter case, it
be overcome by direct compression within the cylindrical mother-cell.
Moreover, in those cases where the adult filament consists of
cylindrical cells, we see that the young, germinating spore, at first
spherical, very soon assumes with growth an elliptical or ovoid form:
the direct result of an incipient anisotropy of its envelope, which
when more developed will convert the ovoid into a cylinder. We may also
notice that a truly cylindrical cell is comparatively rare; for in most
cases, what we call a cylindrical cell shews a distinct bulging of
its sides; it is not truly a cylinder, but a portion of a spheroid or
ellipsoid. {245}
Unicellular organisms in general, including the protozoa, the
unicellular cryptogams, the various bacteria, and the free, isolated
cells, spores, ova, etc. of higher organisms, are referable for the
most part to a very small number of typical forms; but besides a
certain number of others which may be so referable, though obscurely,
there are obviously many others in which either no symmetry is to be
recognized, or in which the form is clearly not one of equilibrium.
Among these latter we have Amoeba itself, and all manner of amoeboid
organisms, and also many curiously shaped cells, such as the
Trypanosomes and various other aberrant Infusoria. We shall return to
the consideration of these; but in the meanwhile it will suffice to
say that, as their surfaces are not equilibrium-surfaces, so neither
are the living cells themselves in any stable equilibrium. On the
contrary, they are in continual flux and movement, each portion of
the surface constantly changing its form, and passing from one phase
to another of an equilibrium which is never stable for more than a
moment. The former class, which rest in stable equilibrium, must fall
(as we have seen) into two classes,—those whose equilibrium arises
from liquid surface-tension alone, and those in whose conformation
some other pressure or restraint has been superimposed upon ordinary
surface-tension.
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