[Illustration: Fig. 80. _Trachelophyllum._ (After Wreszniowski.)]
Precisely the same series of unduloid forms may be traced in even
greater variety among various other families or genera of the
Infusoria. Sometimes, as in Vorticella itself, the unduloid is seen
merely in the contour of the soft semifluid body of the living
animal. At other times, as in Salpingoeca, Tintinnus, and many other
genera, we have a distinct membranous cup, separate from the animal,
but originally secreted by, and moulded upon, its semifluid living
surface. Here we have an excellent illustration of the contrast
between the different ways in which such a structure may be regarded
and interpreted. The teleological explanation is that it is developed
for the sake of protection, as a domicile and shelter for the little
organism within. The mechanical explanation of the physicist (seeking
only after the “efficient,” and not the “final” cause), is that it is
{249} present, and has its actual conformation, by reason of certain
chemico-physical conditions: that it was inevitable, under the given
conditions, that certain constituent substances actually present in
the protoplasm should be aggregated by molecular forces in its surface
layer; that under this adsorptive process, the conditions continuing
favourable, the particles should accumulate and concentrate till
they formed (with the help of the surrounding medium) a pellicle or
membrane, thicker or thinner as the case might be; that this surface
pellicle or membrane was inevitably bound, by molecular forces, to
become a surface of the least possible area which the circumstances
permitted; that in the present case, the symmetry and “freedom” of
the system permitted, and _ipso facto_ caused, this surface to be a
surface of revolution; and that of the few surfaces of revolution
which, as being also surfaces _minimae areae_, were available, the
unduloid was manifestly the one permitted, and _ipso facto_ caused, by
the dimensions of the organisms and other circumstances of the case.
And just as the thickness or thinness of the pellicle was obviously
a subordinate matter, a mere matter of degree, so we also see that
the actual outline of this or that particular unduloid is also a very
subordinate matter, such as physico-chemical variants of a minute kind
would suffice to bring about; for between the various unduloids which
the various species of Vorticella represent, there is no more real
difference than that difference of ratio or degree which exists between
two circles of different diameter, or two lines of unequal length.
{250}
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