Passing to the typical, calcareous-shelled Foraminifera, we have
the most symmetrical of all possible types in the perfect sphere of
Orbulina; this is a pelagic organism, whose floating habitat places it
in a position of perfect symmetry towards all external forces. Save for
one or two other forms which are also spherical, or approximately so,
like Thurammina, the rest of the monothalamic calcareous Foraminifera
are all comprised by naturalists within the genus Lagena. This large
and varied genus consists of “flask-shaped” shells, whose surface is
simply that of an unduloid, or more frequently, like that of a flask
itself, an unduloid combined with a portion of a sphere. We do not know
the circumstances {257} under which the shell of Lagena is formed, nor
the nature of the force by which, during its formation, the surface is
stretched out into the unduloid form; but we may be pretty sure that
it is suspended vertically in the sea, that is to say in a position of
symmetry as regards its vertical axis, about which the unduloid surface
of revolution is symmetrically formed. At the same time we have other
types of the same shell in which the form is more or less flattened;
and these are doubtless the cases in which such symmetry of position
was not present, or was replaced by a broader, lateral contact with the
surface pellicle[305].
[Illustration: Fig. 85. (After Darling.)]
While Orbulina is a simple spherical drop, Lagena suggests to our
minds a “hanging drop,” drawn out to a long and slender neck by
its own weight, aided by the viscosity of the material. Indeed the
various hanging drops, such as Mr C. R. Darling shews us, are the
most beautiful and perfect unduloids, with spherical ends, that it is
possible to conceive. A suitable liquid, a little denser than water
and incapable of mixing with it (such as ethyl benzoate), is poured on
a surface of water. It spreads {258} over the surface and gradually
forms a hanging drop, approximately hemispherical; but as more liquid
is added the drop sinks or rather grows downwards, still adhering
to the surface film; and the balance of forces between gravity and
surface tension results in the unduloid contour, as the increasing
weight of the drop tends to stretch it out and finally break it in
two. At the moment of rupture, by the way, a tiny droplet is formed in
the attenuated neck, such as we described in the normal division of a
cylindrical thread (p. 233).
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