We have innumerable cases, near the tip of a growing filament, where
in like manner the partition-wall which cuts off the terminal {303}
cell constitutes a spherical lens-shaped surface, set normally to the
adjacent walls. At the tips of the branches of many Florideae, for
instance, we find such a lenticular partition. In _Dictyota dichotoma_,
as figured by Reinke, we have a succession of such partitions; and,
by the way, in such cases as these, where the tissues are very
transparent, we have often in optical section a puzzling confusion of
lines; one being the optical section of the curved partition-wall, the
other being the straight linear projection of its outer edge to which
we have already referred. In the conical terminal cell of Chara, we
have the same lens-shaped curve, but a little lower down, where the
sides of the shoot are approximately parallel, we have flat transverse
partitions, at the edges of which, however, we recognise a convexity of
the outer cell-wall and a definite angle of contact, equal on the two
sides of the partition.
[Illustration: Fig. 110. Cells of _Dictyota_. (After Reinke.)]
[Illustration: Fig. 111. Terminal and other cells of _Chara_.]
[Illustration: Fig. 112. Young antheridium of _Chara_.]
In the young antheridia of Chara (Fig. 112), and in the not dissimilar
case of the sporangium (or conidiophore) of Mucor, we easily recognise
the hemispherical form of the septum which shuts off the large
spherical cell from the cylindrical filament. Here, in the first phase
of development, we should have to take into consideration the different
pressures exerted by the single curvature of the cylinder and the
double curvature of its spherical cap (p. 221); and we should find
that the partition would have a somewhat low curvature, with a radius
_less_ than the diameter of the cylinder; which it would have exactly
equalled but for the additional pressure inwards which it receives
{304} from the curvature of the large surrounding sphere. But as the
latter continues to grow, its curvature decreases, and so likewise does
the inward pressure of its surface; and accordingly the little convex
partition bulges out more and more.
――――――――――
In order to epitomise the foregoing facts let the annexed diagrams
(Fig. 113) represent a system of three films, of which one is a
partition-wall between the other two; and let the tensions at the
three surfaces, or the tractions exercised upon a point at their
meeting-place, be proportional to _T_, _T′_ and _t_. Let α, β, γ be, as
in the figure, the opposite angles. Then:
(1) If _T_ be equal to _T′_, and _t_ be relatively insignificant, the
angles α, β will be of 90°.
[Illustration: Fig. 113.]
(2) If _T_ = _T′_, but be a little greater than _t_, then _t_ will
exert an appreciable traction, and α, β will be more than 90°, say, for
instance, 100°.
(3) If _T_ = _T′_ = _t_, then α, β, γ will all equal 120°.
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